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Howell's violet, violette de Howell

cedar glade or glade violet, glade violet

Habit Plants perennial, caulescent, not stoloniferous, 2–44 cm. Plants perennial, acaulescent, not stoloniferous, 5–20 cm; rhizome thick, fleshy.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

Leaves

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

basal, 3, prostrate to ascending;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1.5–7 cm, usually glabrous;

earliest leaf blades ± deltate or 3-lobed, mid-season blades 5–9-lobed, 1–9 × 1–10 cm, base truncate to cuneate, middle lobe lanceolate or spatulate to narrowly ovate, lateral lobes lanceolate or spatulate to falcate, margins serrate, sometimes with deltate or falcate appendages or teeth, ciliate, apex acute, surfaces usually glabrous, rarely pubescent.

Peduncles

2.8–17.8 cm, glabrous or sparsely pubescent.

2–15 cm, usually glabrous, rarely pubescent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 0.5–1 mm;

petals light to dark blue-violet on both surfaces, lower 3 and sometimes upper 2 white basally, lower 3 darker violet-veined, lateral 2 densely bearded, lowest 10–15 mm, spur usually lilac, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on prostrate peduncles.

Capsules

ellipsoid, 7–11 mm, glabrous.

ellipsoid, 11–14 mm, glabrous.

Seeds

light brown, 2.1–2.8 mm.

beige, mottled to bronze, 2–3 mm.

2n

= 40, 80.

= 54.

Viola howellii

Viola egglestonii

Phenology Flowering Apr–Jul. Flowering Mar–May.
Habitat Moist, shady areas, coniferous forests, prairies, along streams Limestone glades and barrens
Elevation 50–1500 m (200–4900 ft) 100–200 m (300–700 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA; IN; KY; TN
[BONAP county map]
Discussion

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

N. H. Russell (1965) was the first to consider Viola egglestonii similar to V. septemloba. After analyzing leaf-blade lobing patterns, L. E. McKinney considered it a subspecies of V. septemloba. Although similarities exist between these taxa, we maintain them as distinct species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 134. FNA vol. 6, p. 129.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. septemloba subsp. egglestonii
Name authority A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) Brainerd: Bull. Torrey Bot. Club 37: 526, plates 34, 35. (1910)
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