Viola howellii |
Viola cuneata |
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Howell's violet, violette de Howell |
northern two-eyed or wedge-leaf violet, northern two-eyed violet, wedge-leaf violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–44 cm. | Plants perennial, caulescent, not stoloniferous, 2–25 cm. |
Stems | 1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome. |
1–3, usually erect, sometimes prostrate or ascending, glabrous, from shallow, fleshy rhizome or deep-seated caudex with fleshy roots. |
Leaves | basal and cauline; basal: 1–6; stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 4–15 cm, glabrous or sparsely puberulent; blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent; cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate; petiole 1–4.5 cm; blade 2–5.1 × 1.2–5.8 cm. |
basal and cauline; basal: 2–6; stipules lanceolate, margins entire or gland-toothed, apex acute; petiole 4.5–9.8 cm, glabrous; blade purple-veined, orbiculate-ovate to deltate, 1–4 × 2.7–3.8 cm, usually shiny, leathery, base cuneate, margins serrate, eciliate, apex acute, mucronulate, surfaces glabrous; cauline similar to basal except: stipules lanceolate to ovate, margins entire or erose to lacerate, with or without gland-tipped processes, apex acute to acuminate; petiole 0.5–20 cm; blade usually rhombic, sometimes orbiculate, 0.9–2.6 × 0.7–1.8 cm, margins crenate to ± serrate, apex acute to obtuse. |
Peduncles | 2.8–17.8 cm, glabrous or sparsely pubescent. |
1–10.5 cm, glabrous. |
Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm; petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep reddish violet abaxially, white adaxially, upper 2 sometimes with purple patch basally, lateral 2 with purple patch basally beyond smaller yellow area, usually bearded, sometimes beardless, lowest purple-veined with yellow area basally, 8–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | ellipsoid, 7–11 mm, glabrous. |
± spherical, 5–9 mm, glabrous. |
Seeds | light brown, 2.1–2.8 mm. |
deep brown-purple, 2.1–3 mm. |
2n | = 40, 80. |
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Viola howellii |
Viola cuneata |
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Phenology | Flowering Apr–Jul. | Flowering Mar–Sep. |
Habitat | Moist, shady areas, coniferous forests, prairies, along streams | Open pine or oak forests, often serpentine soil |
Elevation | 50–1500 m (200–4900 ft) | 600–2200 m (2000–7200 ft) |
Distribution |
CA; OR; WA; BC
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CA; OR
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Discussion | Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola cuneata often occurs on serpentine-derived soil and is closely related to V. flettii, V. hallii, and V. ocellata (V. B. Baird 1942). Originally described as beardless by Watson, the lateral petals of flowers in the same population can be prominently bearded or essentially beardless. The cauline blades of Viola cuneata are often vertical, especially early in season. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 134. | FNA vol. 6, p. 129. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Name authority | A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) | S. Watson: Proc. Amer. Acad. Arts 14: 290. (1879) |
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