Viola howellii |
Viola cucullata |
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Howell's violet, violette de Howell |
blue marsh violet, marsh blue or northern bog or marsh or blue marsh violet, marsh blue violet, violette cucullée |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–44 cm. | Plants perennial, acaulescent, not stoloniferous, 5–20 cm; rhizome thick, fleshy. |
Stems | 1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–6; stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 4–15 cm, glabrous or sparsely puberulent; blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent; cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate; petiole 1–4.5 cm; blade 2–5.1 × 1.2–5.8 cm. |
basal, 4–8, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 6–20 cm, usually glabrous; blade unlobed, ovate to reniform, 1.5–11 × 1–9 cm, base reniform to cordate, margins crenate to serrate, ciliate, apex acute or obtuse to slightly caudate, surfaces glabrous, rarely pubescent. |
Peduncles | 2.8–17.8 cm, glabrous or sparsely pubescent. |
7–25 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm; petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles 3–6 mm; petals light to dark blue-violet to violet on both surfaces, upper 2 and lateral 2 darker basally, lateral 2 densely bearded, lowest white basally, dark purple-veined, 9–13 mm, spur color same as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect peduncles. |
Capsules | ellipsoid, 7–11 mm, glabrous. |
ellipsoid to ± ovoid, 10–15 mm, glabrous. |
Seeds | light brown, 2.1–2.8 mm. |
beige, mottled to bronze, 1–2 mm. |
2n | = 40, 80. |
= 54. |
Viola howellii |
Viola cucullata |
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Phenology | Flowering Apr–Jul. | Flowering Apr–Jun. |
Habitat | Moist, shady areas, coniferous forests, prairies, along streams | Open or wooded wet areas, seeps, springs, swamps, marshes, streamsides |
Elevation | 50–1500 m (200–4900 ft) | 0–2000 m (0–6600 ft) |
Distribution |
CA; OR; WA; BC
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AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; NB; NF; NS; ON; PE; QC; SPM
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Discussion | Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola cucullata reportedly hybridizes with V. sororia (= V. ×bissellii House), V. brittoniana (= V. ×notabilis E. P. Bicknell), and V. sagittata var. sagittata (= V. ×porteriana House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 134. | FNA vol. 6, p. 128. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. cucullata var. leptosepala, V. cucullata var. macrotis, V. cucullata var. microtitis, V. dicksonii, V. leptosepala, V. macrotis | |
Name authority | A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887) | Aiton: Hort. Kew. 3: 288. (1789) |
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