Flora of North America species comparison

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Viola hirsutula

Viola sempervirens

southern woodland violet

evergreen violet, redwood violet, redwoods violet, trailing yellow violet, violette toujours verte
Habit Plants perennial, acaulescent, not stoloniferous, 2–15 cm; rhizome thick, fleshy. Plants perennial, caulescent, stoloniferous, 10–30 cm; stolons green or reddish, leafy, sometimes rooting at nodes, becoming lignified in age.
Stems

1–5, prostrate, spreading, glabrous or sparsely puberulent, from current and/or previous year’s growth, on usually vertical, fleshy rhizome, rooting and forming rosettes at or near tip; rooted rosettes often develop into an erect, fleshy caudex from which new stems are produced.
Leaves

basal, 2–4, prostrate or nearly so;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1–10 cm, usually glabrous;

blade unlobed, reniform to ovate, 1–6 × 1–5 cm, base cordate, margins serrate, ciliate or eciliate, apex rounded, adaxial leaf surface with silvery strigose patches.

evergreen, basal and cauline;

basal: 1–6(–10);

stipules deltate to ovate or linear-lanceolate, margins entire or glandular-toothed, apex acute to long-acuminate;

petiole 2–16 cm, glabrous;

blade often purple-spotted abaxially and/or adaxially, orbiculate to ovate, 1–4.5 × 2–3.9 cm, base cordate to truncate, margins crenate, eciliate, apex blunt to obtuse, mucronulate, surfaces glabrous or with scattered bristles on one or both surfaces;

cauline similar to basal except: stipules deltate to lanceolate, margins entire or sparingly toothed;

petiole 0.3–3 cm;

blade 1.2–2.2 × 1.2–2 cm.
Peduncles

2–12 cm, glabrous.

5–10 cm, glabrous.
Flowers

sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;

petals light to deep reddish violet to blue-violet on both surfaces, 3 lower whitish basally, dark violet-veined, lateral 2 densely bearded, lowest 11–17 mm, spur violet, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

sepals lanceolate to ovate-lanceolate, margins eciliate, auricles 1–2 mm;

petals lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 8–17 mm, spur yellow or whitish, gibbous, 1–2.5 mm;

style head bearded; cleistogamous flowers axillary.
Capsules

ellipsoid, 8–12 mm, glabrous.

mottled with purple, spherical to ovoid, 5–8 mm, glabrous.
Seeds

beige, mottled to bronze, 1–1.5 mm.

brown, tinged purple, 2–2.5 mm.
2n

= 54.

= 24, 48.

Viola hirsutula

Viola sempervirens
Phenology Flowering Mar–Jun. Flowering Jan–Jul.
Habitat Dry to mesic deciduous woodlands Redwood forests, other coastal forests, Douglas fir, other coniferous forests
Elevation 100–1000 m (300–3300 ft) 30–1400 m (100–4600 ft)
Distribution
from FNA
AL; CT; DC; DE; FL; GA; IN; KY; MD; MS; NC; NJ; NY; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; ID; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola hirsutula exhibits little phenotypic plasticity. Although considered by N. L. Gil-Ad (1997) to be a putative hybrid, he did not identify the potential parents and chose not to recognize it. A. Cronquist in H. A. Gleason and Cronquist (1991) considered V. hirsutula a form of V. villosa. Viola hirsutula is considered one of the more distinct species throughout its range by McKinney because of the pattern of silvery strigose patches on adaxial leaf surfaces and a more prostrate growth habit than similar species. He believes V. hirsutula is one of the least variable species in the flora. Viola hirsutula appears to occur much less frequently than herbarium collections suggest. Some herbarium specimens are misidentified and represent depauperate plants, usually of V. sororia. These specimens are often hairy on the adaxial leaf surfaces but lack the silvery patches.

Viola hirsutula reportedly hybridizes with V. sororia var. sororia (= V. ×cordifolia [Nuttall] Schweinitz) and V. sagittata var. sagittata (= V. ×redacta House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In California, Viola sempervirens occurs in shaded redwood forests and other coastal forest habitats. In Oregon and Washington, it occurs in Douglas fir and other coniferous forests, where it can form mats (clones) one meter or more in diameter; its prostrate, spreading growth habit is similar to V. walteri. The leafy stems of V. sempervirens are similar to the leafy stolons of V. odorata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 6, p. 133. FNA vol. 6, p. 157.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. sarmentosa, V. sempervirens subsp. orbiculoides
Name authority Brainerd: Rhodora 9: 98. (1907) Greene: Pittonia 4: 8. (1899)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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