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Viola hallii

Hall's violet, Oregon violet, wild pansy

sagebrush violet, valley violet, valley yellow violet, yellow prairie violet, yellow sagebrush violet

Habit Plants perennial, caulescent, not stoloniferous, 5–22 cm. Plants perennial, caulescent, not stoloniferous, 2–18 cm.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous, clustered on single, short, vertical, deep-seated caudex.

1–5, decumbent or ascending to erect, leafy proximally and distally, ca. 1/2 subterranean, glabrate to puberulent, on caudex from usually vertical, subligneous rhizome.

Leaves

basal and cauline;

basal: 1–4, palmately compound, ± 2-ternate or 3-ternate, leaflets 3;

stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute;

petiole 5–8 cm, glabrous;

blade ovate to deltate, 2.8–6 × 2.6–6.5 cm, ± coriaceous, base tapered, ultimate lobes narrowly elliptic, lanceolate, or oblanceolate, 1–7 mm wide, margins entire, ciliate or eciliate, apex acute, mucronulate, surfaces glabrous;

cauline similar to basal except: stipules usually lanceolate, sometimes broadly ovate, ± leaflike, margins toothed;

petiole 1.3–6 cm;

blade 2–4.8 × 1.2–5.5 cm.

basal and cauline;

basal: 1–6;

stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, sometimes divided into filiform processes;

petiole 3–10 cm, glabrous or puberulent;

blade ovate to oblong-ovate, 1.5–4.3 × 0.9–1.1 cm, base usually truncate, sometimes attenuate, margins entire or serrulate, sometimes sinuate, ciliate (sometimes only on proximal 1/2 of leaf), apex acute to obtuse, surfaces glabrous or puberulent;

cauline similar to basal except: stipules linear to linear-lanceolate, margins ± entire, apex acute;

petiole 1.5–9.2 cm;

blade ovate or lanceolate to elliptic, 2.3–4 × 1–2 cm, length ca. 2.2 times width, base usually ± truncate to subcordate, sometimes attenuate on new leaves, margins sinuate, apex acute.

Peduncles

2.5–11 cm, glabrous.

3–11.5 cm, glabrous or puberulent.

Flowers

sepals lanceolate to ovate, margins ciliate, auricles 0.5–1 mm;

petals: upper 2 almost black abaxially, dark reddish violet adaxially, lower 3 pale yellow, cream, or ± white, lateral 2 bearded, with deep yellow to orange patch basally, dark reddish violet-veined, lowest with deep yellow to orange patch basally, dark reddish violet-veined, 5–18 mm, spur yellow, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate, margins eciliate, auricles 0.5–1 mm;

petals deep lemon-yellow adaxially or on both surfaces, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 9–14 mm, spur yellow, gibbous, 0.8–1.2 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 4–12 mm, glabrous.

spherical, ca. 5 mm, glabrous or finely puberulent.

Seeds

light brown, shiny, 3.2–3.5 mm.

tan, 2.1–2.2 mm, elaiosome extending beyond and covering funiculus.

2n

= 60, 72.

= 12.

Viola hallii

Viola vallicola

Phenology Flowering Apr–Jul. Flowering late Mar–early Jul.
Habitat Open forests, grassy hills, chaparral, frequently serpentine or gravelly soil Sagebrush flats, prairie grasslands, open forests, juniper woodlands
Elevation 500–2100 m (1600–6900 ft) 400–2800 m (1300–9200 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; KS; MT; ND; SD; WY; BC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola hallii was discovered on the grounds of Willamette University in Salem, Oregon, by Elihu Hall, a professor at that institution (V. B. Baird 1942). Leaves of V. hallii are similar to V. beckwithii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. M. Fabijan et al. (1987) distinguished two varieties of Viola vallicola based on geographic location and type of leaf flavonoids: 1) var. major (Hooker) Fabijan, occurring west of the Continental Divide with leaf flavonoids primarily kaempferol derivatives; and 2) var. vallicola occurring east of the Continental Divide with leaf flavonoids all apigenin derivatives. Fabijan et al. reported that var. major (valley violet) occurs in sagebrush flats, prairie grasslands, open forests, juniper woodlands, 400–2800 m, in Alberta, Colorado, Idaho, Montana, Nevada, Oregon, Utah, and Washington.

M. S. Baker (1957) noted that there is only a tendency for leaves of Viola vallicola to be wide with truncate bases; on some plants only a few leaves are truncate. Images of type specimens at NY show truncate leaf bases for most basal and cauline leaves, with only some cauline leaves being attenuate. The key to V. vallicola in D. M. Fabijan et al. (1987) described the cauline leaf bases as truncate with some later cauline leaves becoming cuneate. G. Davidse (1976) stated that the V. vallicola plants he studied had cordate to truncate leaf bases; he made no distinction between basal and cauline leaves. Baker stated that V. vallicola may have given rise to V. nuttallii through a doubling of its chromosomes.

Because intermediate leaf forms are found in areas where Viola vallicola and V. nuttallii are sympatric, some question the specific status of V. vallicola. No hybrids involving V. nuttallii and V. vallicola are known (D. M. Fabijan et al. 1987).

V. Harms (pers. comm.) reported that Viola vallicola is frequent in southern Saskatchewan grasslands, usually occurring with V. nuttallii and appearing the more frequent of the two.

Observed pollinators of Viola vallicola in the intermountain region include flies in the genera Bombylius Linnaeus and Eristalis Latreille (G. Davidse 1976).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 133. FNA vol. 6, p. 162.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. villosa, V. walteri
Synonyms Crocion vallicola, V. nuttallii subsp. vallicola, V. nuttallii var. vallicola, V. physalodes, V. russellii, V. subsagittifolia
Name authority A. Gray: Proc. Amer. Acad. Arts 8: 377. (1872) A. Nelson: Bull. Torrey Bot. Club 26: 128. (1899)
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