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Viola hallii

Hall's violet, Oregon violet, wild pansy

Astoria violet, Canary violet, marsh violet, upland yellow violet, yellow montane violet

Habit Plants perennial, caulescent, not stoloniferous, 5–22 cm. Plants perennial, caulescent, not stoloniferous, 5.5–36.5 cm.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous, clustered on single, short, vertical, deep-seated caudex.

1–3(–5), prostrate, decumbent, or erect, leafy proximally and distally, glabrous or puberulent, on caudex from usually vertical, subligneous rhizome.

Leaves

basal and cauline;

basal: 1–4, palmately compound, ± 2-ternate or 3-ternate, leaflets 3;

stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute;

petiole 5–8 cm, glabrous;

blade ovate to deltate, 2.8–6 × 2.6–6.5 cm, ± coriaceous, base tapered, ultimate lobes narrowly elliptic, lanceolate, or oblanceolate, 1–7 mm wide, margins entire, ciliate or eciliate, apex acute, mucronulate, surfaces glabrous;

cauline similar to basal except: stipules usually lanceolate, sometimes broadly ovate, ± leaflike, margins toothed;

petiole 1.3–6 cm;

blade 2–4.8 × 1.2–5.5 cm.

basal and cauline;

basal: 1–5;

stipules adnate to proximal 1/3 of petiole, forming 2 narrow, linear-lanceolate wings, margins entire, apex of each wing free, acute;

petiole 2.6–19.2 cm, glabrous or densely puberulent;

blade usually elliptic to ovate, sometimes oblong-lanceolate to ± orbiculate, 1.7–14(–17) × 0.8–5.3(–6.7) cm, base attenuate to ± truncate or subcordate, often oblique, margins usually crenate, serrulate, or serrate, sometimes entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or densely puberulent;

cauline similar to basal except: stipules lanceolate to ovate, margins entire or toothed, with or without gland-tipped projections, apex acute to acuminate;

petiole 1.3–16.2 cm, glabrous or puberulent;

blade 2.3–11(–14.8) × 1.1–3.6(–5.5) cm, length 1.1–6.5 times width.

Peduncles

2.5–11 cm, glabrous.

4.4–27 cm, glabrous or puberulent.

Flowers

sepals lanceolate to ovate, margins ciliate, auricles 0.5–1 mm;

petals: upper 2 almost black abaxially, dark reddish violet adaxially, lower 3 pale yellow, cream, or ± white, lateral 2 bearded, with deep yellow to orange patch basally, dark reddish violet-veined, lowest with deep yellow to orange patch basally, dark reddish violet-veined, 5–18 mm, spur yellow, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate, margins eciliate, auricles 1–2 mm;

petals deep lemon-yellow adaxially, upper 2, and sometimes lateral 2, brownish purple abaxially, lower 3 brownish purple-veined, lateral 2 sparsely bearded, lowest 11–19 mm, spur yellow or pale green, gibbous, 0.5–3 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ellipsoid, 4–12 mm, glabrous.

ellipsoid to oblong, 6–14 mm, glabrous or finely puberulent.

Seeds

light brown, shiny, 3.2–3.5 mm.

medium to dark brown or red-brown, 2–3 mm, elaiosome completely covering funiculus.

2n

= 60, 72.

= 36, 48.

Viola hallii

Viola praemorsa

Phenology Flowering Apr–Jul.
Habitat Open forests, grassy hills, chaparral, frequently serpentine or gravelly soil
Elevation 500–2100 m (1600–6900 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola hallii was discovered on the grounds of Willamette University in Salem, Oregon, by Elihu Hall, a professor at that institution (V. B. Baird 1942). Leaves of V. hallii are similar to V. beckwithii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Viola praemorsa is a complex group that appears to be evolving. Sympatric populations and the similarity of their flowers provide opportunities for hybridization and introgression (D. M. Fabijan et al. 1987). Some botanists have suggested that all taxa in the V. nuttallii complex, which includes V. bakeri, V. nuttallii, V. praemorsa, V. tomentosa, and V. vallicola, should be treated as varieties of V. nuttallii, as C. L. Hitchcock et al. (1955–1969, vol. 3) did, on the basis that these taxa are more or less sympatric in range, intergrade with one another, and possess no distinctive gross morphological features by which they can be consistently recognized. S. L. Welsh et al. (1987) treated V. praemorsa and V. linguaefolia Nuttall as synonymous with V. nuttallii, stating that V. nuttallii (and related taxa) are acaulescent to short-caulescent. Although some plants have short internodes, all plants in the V. nuttallii complex are caulescent. We acknowledge the complexity and taxonomic difficulties inherent in the V. nuttallii complex; patterns of morphology, cytology, and leaf flavonoid chemistry provide a basis for recognizing infrataxa. Fabijan et al. conducted the most extensive study of the complex to date; their treatment of V. praemorsa is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal and cauline leaf blade bases ± truncate to subcordate; basal leaf blades 1.7–5.9(–7.3) cm, length 1.1–3 times width; plants 5.5–21.5 cm.
var. praemorsa
1. Basal and cauline leaf blade bases attenuate; basal leaf blades 2.7–17 cm, length 1.7–6.5 times width; plants 10.5–36.5 cm
→ 2
2. Basal leaf blades (3.4–)5.3–17 cm, usually ovate to elliptic, sometimes oblong-lanceolate, length 1.8–6.5 times width; peduncles 5.9–27 cm.
var. flavovirens
2. Basal leaf blades 2.7–8.5 cm, usually ovate to elliptic, sometimes oblong-lanceolate, length 1.7–3.4 times width; peduncles 5–15 cm.
var. linguifolia
Source FNA vol. 6, p. 133. FNA vol. 6, p. 145.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. praemorsa var. flavovirens, V. praemorsa var. linguifolia, V. praemorsa var. praemorsa
Synonyms V. nuttallii subsp. praemorsa, V. nuttallii var. praemorsa
Name authority A. Gray: Proc. Amer. Acad. Arts 8: 377. (1872) Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1254. (1829)
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