Viola hallii |
Viola nephrophylla |
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Hall's violet, Oregon violet, wild pansy |
kidney-leaf violet, Leconte violet, northern bog violet, violette néphrophylle |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–22 cm. | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous, clustered on single, short, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–4, palmately compound, ± 2-ternate or 3-ternate, leaflets 3; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 5–8 cm, glabrous; blade ovate to deltate, 2.8–6 × 2.6–6.5 cm, ± coriaceous, base tapered, ultimate lobes narrowly elliptic, lanceolate, or oblanceolate, 1–7 mm wide, margins entire, ciliate or eciliate, apex acute, mucronulate, surfaces glabrous; cauline similar to basal except: stipules usually lanceolate, sometimes broadly ovate, ± leaflike, margins toothed; petiole 1.3–6 cm; blade 2–4.8 × 1.2–5.5 cm. |
basal, 4–7, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–25 cm, usually glabrous; blade usually grayish green or purplish green abaxially, unlobed, ovate, reniform, or broadly reniform to orbiculate, 1–7 × 1–7 cm, somewhat fleshy, base broadly cordate or reniform to ± truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually glabrous, rarely sparsely pubescent. |
Peduncles | 2.5–11 cm, glabrous. |
3–25 cm, usually glabrous. |
Flowers | sepals lanceolate to ovate, margins ciliate, auricles 0.5–1 mm; petals: upper 2 almost black abaxially, dark reddish violet adaxially, lower 3 pale yellow, cream, or ± white, lateral 2 bearded, with deep yellow to orange patch basally, dark reddish violet-veined, lowest with deep yellow to orange patch basally, dark reddish violet-veined, 5–18 mm, spur yellow, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals ovate, margins usually eciliate, auricles 1–2 mm; petals deep bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, upper 2 sometimes sparsely bearded, lowest densely bearded or beardless, 10–28 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on erect to ascending peduncles. |
Capsules | ellipsoid, 4–12 mm, glabrous. |
ovoid, 5–10 mm, glabrous. |
Seeds | light brown, shiny, 3.2–3.5 mm. |
beige to brown or dark brown, 1.5–2.5 mm. |
2n | = 60, 72. |
= 54. |
Viola hallii |
Viola nephrophylla |
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Phenology | Flowering Apr–Jul. | Flowering Jan–Sep. |
Habitat | Open forests, grassy hills, chaparral, frequently serpentine or gravelly soil | Wet habitats in saturated soil in prairies, pastures, bogs, fens, sedge meadows, rocky shores of lakes and streams, limestone outcrops, gravelly calcareous stream beds |
Elevation | 500–2100 m (1600–6900 ft) | 100–3000 m (300–9800 ft) |
Distribution |
CA; OR
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AR; AZ; CA; CO; CT; IA; ID; IL; KS; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OH; OR; PA; SD; TX; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; PE; QC; SK; YT
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Discussion | Viola hallii was discovered on the grounds of Willamette University in Salem, Oregon, by Elihu Hall, a professor at that institution (V. B. Baird 1942). Leaves of V. hallii are similar to V. beckwithii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola nephrophylla has the widest distribution in North America of any of the acaulescent blue violets. There is some question whether the species we now know as Viola nephrophylla fits with the protologue and type specimen(s). McKinney examined the type designated by Greene; it appears that what most botanists have called V. nephrophylla may be an undescribed taxon. N. L. Gil-Ad (1997) believed Greene’s type contained plants that were genetically impure based on seed coat micromorphology and suggested the specimens might be a mixture of both orthospecies and introgressants. Viola mccabeiana was described by M. S. Baker in 1940 from collections made in British Columbia. Later, Baker (1949b) discussed the reasons why the name V. mccabeiana needed to be abandoned. C. L. Hitchcock et al. (1955–1969, vol. 3) treated V. mccabeiana as synonymous with V. nephrophylla var. cognata. While Viola mccabeiana is recognized in Canada (L. Brouillet et al., http://data.canadensys.net/vascan/), we believe it is best included within V. nephrophylla. Viola nephrophylla reportedly hybridizes with V. cucullata (= V. ×insessa House), V. sororia var. sororia (= V. ×napae House), and V. affinis (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 133. | FNA vol. 6, p. 139. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. arizonica, V. austiniae, V. cognata, V. crassula, V. galacifolia, V. lunellii, V. mccabeiana, V. nephrophylla var. arizonica, V. nephrophylla var. cognata, V. peramoena, V. pratincola, V. prionosepala, V. retusa, V. subjuncta, V. vagula | |
Name authority | A. Gray: Proc. Amer. Acad. Arts 8: 377. (1872) | Greene: Pittonia 3: 144. (1896) |
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