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pioneer violet, smooth yellow violet, stream or smooth yellow or yellow wood or wood or smooth yellow woodland violet, stream violet, violette glabre, yellow wood violet

Astoria violet, Canary violet, marsh violet, upland yellow violet, yellow montane violet

Habit Plants perennial, caulescent, not stoloniferous, 3–38 cm. Plants perennial, caulescent, not stoloniferous, 5.5–36.5 cm.
Stems

1–3, erect or prostrate, leafless proximally, leafy distally, glabrous or finely puberulent, on caudex from fleshy rhizome.

1–3(–5), prostrate, decumbent, or erect, leafy proximally and distally, glabrous or puberulent, on caudex from usually vertical, subligneous rhizome.

Leaves

basal and cauline;

basal: 0–4(–7);

stipules ovate to obovate, margins entire, crenate, or serrate, usually glandular, apex acute to obtuse;

petiole 7–27.5 cm, glabrous or puberulent;

blade usually reniform to ovate, sometimes orbiculate, 3.3–8.5 × 2–9.3 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or finely puberulent;

cauline similar to basal except: usually restricted to distal ends of naked stems;

stipules ovate to oblong, margins erose or subserrate, often glandular, apex acute to acuminate;

petiole 0.2–2.9 cm, glabrous or finely puberulent;

blade ovate to deltate, 1.4–5.7 × 0.8–4.7 cm, base cordate to truncate, margins crenate to ± serrate, ciliate (sometimes limited to proximal half), apex acute.

basal and cauline;

basal: 1–5;

stipules adnate to proximal 1/3 of petiole, forming 2 narrow, linear-lanceolate wings, margins entire, apex of each wing free, acute;

petiole 2.6–19.2 cm, glabrous or densely puberulent;

blade usually elliptic to ovate, sometimes oblong-lanceolate to ± orbiculate, 1.7–14(–17) × 0.8–5.3(–6.7) cm, base attenuate to ± truncate or subcordate, often oblique, margins usually crenate, serrulate, or serrate, sometimes entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or densely puberulent;

cauline similar to basal except: stipules lanceolate to ovate, margins entire or toothed, with or without gland-tipped projections, apex acute to acuminate;

petiole 1.3–16.2 cm, glabrous or puberulent;

blade 2.3–11(–14.8) × 1.1–3.6(–5.5) cm, length 1.1–6.5 times width.

Peduncles

2–8 cm, glabrous or pubescent.

4.4–27 cm, glabrous or puberulent.

Flowers

sepals linear-lanceolate, margins eciliate, auricles 0.5–1 mm;

petals deep lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 6–18 mm, spur yellow to greenish, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate, margins eciliate, auricles 1–2 mm;

petals deep lemon-yellow adaxially, upper 2, and sometimes lateral 2, brownish purple abaxially, lower 3 brownish purple-veined, lateral 2 sparsely bearded, lowest 11–19 mm, spur yellow or pale green, gibbous, 0.5–3 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ovoid to ellipsoid, 7–13 mm, glabrous.

ellipsoid to oblong, 6–14 mm, glabrous or finely puberulent.

Seeds

light to dark brown, shiny, 2–2.2 mm.

medium to dark brown or red-brown, 2–3 mm, elaiosome completely covering funiculus.

2n

= 24.

= 36, 48.

Viola glabella

Viola praemorsa

Phenology Flowering Mar–Aug.
Habitat Damp, wet, or shady places in forests, stream banks
Elevation 0–2600 m (0–8500 ft)
Distribution
from FNA
AK; CA; ID; MT; OR; WA; AB; BC; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola glabella is similar in overall appearance to V. pubescens. The cauline leaves are sometimes described as appearing only near the apex of the stem (P. A. Munz 1959), but often a leaf occurs from a node on the stem below the apex.

Viola californica was described from collections made in Humboldt and Trinity counties, California, and may have arisen as a hybrid between V. glabella and V. lobata var. integrifolia. It is apparently limited to Abies concolor forests above 1520 m (M. S. Baker 1953). Study of V. californica is needed to determine if taxonomic recognition is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Viola praemorsa is a complex group that appears to be evolving. Sympatric populations and the similarity of their flowers provide opportunities for hybridization and introgression (D. M. Fabijan et al. 1987). Some botanists have suggested that all taxa in the V. nuttallii complex, which includes V. bakeri, V. nuttallii, V. praemorsa, V. tomentosa, and V. vallicola, should be treated as varieties of V. nuttallii, as C. L. Hitchcock et al. (1955–1969, vol. 3) did, on the basis that these taxa are more or less sympatric in range, intergrade with one another, and possess no distinctive gross morphological features by which they can be consistently recognized. S. L. Welsh et al. (1987) treated V. praemorsa and V. linguaefolia Nuttall as synonymous with V. nuttallii, stating that V. nuttallii (and related taxa) are acaulescent to short-caulescent. Although some plants have short internodes, all plants in the V. nuttallii complex are caulescent. We acknowledge the complexity and taxonomic difficulties inherent in the V. nuttallii complex; patterns of morphology, cytology, and leaf flavonoid chemistry provide a basis for recognizing infrataxa. Fabijan et al. conducted the most extensive study of the complex to date; their treatment of V. praemorsa is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal and cauline leaf blade bases ± truncate to subcordate; basal leaf blades 1.7–5.9(–7.3) cm, length 1.1–3 times width; plants 5.5–21.5 cm.
var. praemorsa
1. Basal and cauline leaf blade bases attenuate; basal leaf blades 2.7–17 cm, length 1.7–6.5 times width; plants 10.5–36.5 cm
→ 2
2. Basal leaf blades (3.4–)5.3–17 cm, usually ovate to elliptic, sometimes oblong-lanceolate, length 1.8–6.5 times width; peduncles 5.9–27 cm.
var. flavovirens
2. Basal leaf blades 2.7–8.5 cm, usually ovate to elliptic, sometimes oblong-lanceolate, length 1.7–3.4 times width; peduncles 5–15 cm.
var. linguifolia
Source FNA vol. 6, p. 131. FNA vol. 6, p. 145.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. praemorsa var. flavovirens, V. praemorsa var. linguifolia, V. praemorsa var. praemorsa
Synonyms V. californica, V. canadensis var. sitchensis, V. glabella var. remotifolia V. nuttallii subsp. praemorsa, V. nuttallii var. praemorsa
Name authority Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 142. (1838) Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1254. (1829)
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