Viola glabella |
Viola pedunculata |
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pioneer violet, smooth yellow violet, stream or smooth yellow or yellow wood or wood or smooth yellow woodland violet, stream violet, violette glabre, yellow wood violet |
California golden violet, johnny-jump-up, wild pansy, yellow pansy |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–38 cm. | Plants perennial, caulescent, not stoloniferous, 5–39 cm. |
Stems | 1–3, erect or prostrate, leafless proximally, leafy distally, glabrous or finely puberulent, on caudex from fleshy rhizome. |
1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots. |
Leaves | basal and cauline; basal: 0–4(–7); stipules ovate to obovate, margins entire, crenate, or serrate, usually glandular, apex acute to obtuse; petiole 7–27.5 cm, glabrous or puberulent; blade usually reniform to ovate, sometimes orbiculate, 3.3–8.5 × 2–9.3 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or finely puberulent; cauline similar to basal except: usually restricted to distal ends of naked stems; stipules ovate to oblong, margins erose or subserrate, often glandular, apex acute to acuminate; petiole 0.2–2.9 cm, glabrous or finely puberulent; blade ovate to deltate, 1.4–5.7 × 0.8–4.7 cm, base cordate to truncate, margins crenate to ± serrate, ciliate (sometimes limited to proximal half), apex acute. |
cauline; stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate; petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate; blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent. |
Peduncles | 2–8 cm, glabrous or pubescent. |
2.9–20 cm, sparsely to densely puberulent. |
Flowers | sepals linear-lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 6–18 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm; petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm; style head bearded; cleistogamous flowers absent. |
Capsules | ovoid to ellipsoid, 7–13 mm, glabrous. |
ellipsoid, 5–11 mm, glabrous. |
Seeds | light to dark brown, shiny, 2–2.2 mm. |
dark brown or black, shiny, 2.7 mm. |
2n | = 24. |
= 12. |
Viola glabella |
Viola pedunculata |
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Phenology | Flowering Mar–Aug. | Flowering Feb–Apr. |
Habitat | Damp, wet, or shady places in forests, stream banks | Open, grassy coastal and inland slopes and hillsides, usually in full sun, chaparral, foothill and oak woodland |
Elevation | 0–2600 m (0–8500 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AK; CA; ID; MT; OR; WA; AB; BC; Asia
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CA; Mexico (Baja California)
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Discussion | Viola glabella is similar in overall appearance to V. pubescens. The cauline leaves are sometimes described as appearing only near the apex of the stem (P. A. Munz 1959), but often a leaf occurs from a node on the stem below the apex. Viola californica was described from collections made in Humboldt and Trinity counties, California, and may have arisen as a hybrid between V. glabella and V. lobata var. integrifolia. It is apparently limited to Abies concolor forests above 1520 m (M. S. Baker 1953). Study of V. californica is needed to determine if taxonomic recognition is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex. Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata. Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 144. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. californica, V. canadensis var. sitchensis, V. glabella var. remotifolia | V. pedunculata subsp. tenuifolia |
Name authority | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 142. (1838) | Torrey & A. Gray: Fl. N. Amer. 1: 141. (1838) |
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