Viola glabella |
Viola blanda |
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pioneer violet, smooth yellow violet, stream or smooth yellow or yellow wood or wood or smooth yellow woodland violet, stream violet, violette glabre, yellow wood violet |
sweet white violet, violette agréable |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–38 cm. | Plants perennial, acaulescent, stoloniferous, 3–20 cm; stolons pale, often rooting and leafy at nodes; rhizome short, slender, fleshy. |
Stems | 1–3, erect or prostrate, leafless proximally, leafy distally, glabrous or finely puberulent, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 0–4(–7); stipules ovate to obovate, margins entire, crenate, or serrate, usually glandular, apex acute to obtuse; petiole 7–27.5 cm, glabrous or puberulent; blade usually reniform to ovate, sometimes orbiculate, 3.3–8.5 × 2–9.3 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or finely puberulent; cauline similar to basal except: usually restricted to distal ends of naked stems; stipules ovate to oblong, margins erose or subserrate, often glandular, apex acute to acuminate; petiole 0.2–2.9 cm, glabrous or finely puberulent; blade ovate to deltate, 1.4–5.7 × 0.8–4.7 cm, base cordate to truncate, margins crenate to ± serrate, ciliate (sometimes limited to proximal half), apex acute. |
basal, 2–9, prostrate to ascending; stipules linear-lanceolate, margins entire, apex acute; petiole 2–11 cm, usually sparsely pubescent; blade unlobed, reniform to ovate, 2–4 × 2–4 cm, base cordate, lobes often overlapping, margins serrate, ciliate or eciliate, apex rounded or acute to acuminate, surfaces sometimes glabrous, usually sparsely pubescent adaxially. |
Peduncles | 2–8 cm, glabrous or pubescent. |
3–11 cm, glabrous or pubescent. |
Flowers | sepals linear-lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 6–18 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins mostly eciliate, auricles 1–2 mm; petals white on both surfaces, lower 3 purple-veined, lateral 2 usually beardless, lowest 8–10 mm, spur white, gibbous, 1–2 mm; style head beardless; cleistogamous flowers axillary. |
Capsules | ovoid to ellipsoid, 7–13 mm, glabrous. |
ovoid to ellipsoid, 4–6 mm, glabrous. |
Seeds | light to dark brown, shiny, 2–2.2 mm. |
beige to bronze, 1.5–2 mm. |
2n | = 24. |
= 44, 48. |
Viola glabella |
Viola blanda |
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Phenology | Flowering Mar–Aug. | Flowering Apr–Jun. |
Habitat | Damp, wet, or shady places in forests, stream banks | Rich woods |
Elevation | 0–2600 m (0–8500 ft) | 30–2000 m (100–6600 ft) |
Distribution |
AK; CA; ID; MT; OR; WA; AB; BC; Asia
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AL; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; NC; ND; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; AB; MB; NB; NL; NS; ON; PE; QC; SK
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Discussion | Viola glabella is similar in overall appearance to V. pubescens. The cauline leaves are sometimes described as appearing only near the apex of the stem (P. A. Munz 1959), but often a leaf occurs from a node on the stem below the apex. Viola californica was described from collections made in Humboldt and Trinity counties, California, and may have arisen as a hybrid between V. glabella and V. lobata var. integrifolia. It is apparently limited to Abies concolor forests above 1520 m (M. S. Baker 1953). Study of V. californica is needed to determine if taxonomic recognition is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola blanda occurs in small colonies; individual plants are interconnected by stolons. Whether to recognize Viola incognita at any taxonomic level is currently unresolved. It is said to have pubescent leaf blades, greenish peduncles, nontwisted lateral petals, and a preference for moister habitats. Most of these characters fall within the range of variation observed in V. blanda. N. H. Russell (1965) noted that Viola incognita is principally found in glaciated areas whereas V. blanda is found in nonglaciated areas. Á. Löve and D. Löve (1982b) and J. M. Canne (1987) reported a chromosome count of 2n = 44; J. Clausen (1929) and A. Gershoy (1934) reported 2n = 48. V. B. Baird (1942) reported that V. blanda (and V. incognita) have fragrant flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 124. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. californica, V. canadensis var. sitchensis, V. glabella var. remotifolia | V. blanda var. palustriformis, V. incognita, V. incognita var. forbesii, V. leconteana |
Name authority | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 142. (1838) | Willdenow: Hort. Berol. 1(2): plate 24. (1804) |
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