Viola glabella |
Viola affinis |
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pioneer violet, smooth yellow violet, stream or smooth yellow or yellow wood or wood or smooth yellow woodland violet, stream violet, violette glabre, yellow wood violet |
Le Conte's violet, sand violet, violette affine |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–38 cm. | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. |
Stems | 1–3, erect or prostrate, leafless proximally, leafy distally, glabrous or finely puberulent, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 0–4(–7); stipules ovate to obovate, margins entire, crenate, or serrate, usually glandular, apex acute to obtuse; petiole 7–27.5 cm, glabrous or puberulent; blade usually reniform to ovate, sometimes orbiculate, 3.3–8.5 × 2–9.3 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or finely puberulent; cauline similar to basal except: usually restricted to distal ends of naked stems; stipules ovate to oblong, margins erose or subserrate, often glandular, apex acute to acuminate; petiole 0.2–2.9 cm, glabrous or finely puberulent; blade ovate to deltate, 1.4–5.7 × 0.8–4.7 cm, base cordate to truncate, margins crenate to ± serrate, ciliate (sometimes limited to proximal half), apex acute. |
basal, 1–6, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–10 cm, glabrous; blade green abaxially, unlobed, narrowly to broadly ovate or narrowly deltate, 1.5–10 × 1.5–10 cm, not fleshy, base cordate to broadly cordate or almost truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, surfaces sparsely pubescent adaxially, rarely glabrous. |
Peduncles | 2–8 cm, glabrous or pubescent. |
3–15 cm, glabrous or pubescent. |
Flowers | sepals linear-lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 6–18 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals lavender-violet to dull reddish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, lowest 10–22 mm, usually obviously bearded, rarely beardless, spur white or same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers from prostrate to ascending peduncles. |
Capsules | ovoid to ellipsoid, 7–13 mm, glabrous. |
often reddish or purplish-flecked or green, ellipsoid, 5–10 mm, glabrous or puberulent. |
Seeds | light to dark brown, shiny, 2–2.2 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 24. |
= 54. |
Viola glabella |
Viola affinis |
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Phenology | Flowering Mar–Aug. | Flowering Apr–Jun. |
Habitat | Damp, wet, or shady places in forests, stream banks | Open or wooded wet areas, meadows, stream banks, thickets, shores of lakes, seasonally dry areas. |
Elevation | 0–2600 m (0–8500 ft) | 100–2000 m (300–6600 ft) |
Distribution |
AK; CA; ID; MT; OR; WA; AB; BC; Asia
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CT; DC; DE; IL; IN; MA; MD; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; ON; QC |
Discussion | Viola glabella is similar in overall appearance to V. pubescens. The cauline leaves are sometimes described as appearing only near the apex of the stem (P. A. Munz 1959), but often a leaf occurs from a node on the stem below the apex. Viola californica was described from collections made in Humboldt and Trinity counties, California, and may have arisen as a hybrid between V. glabella and V. lobata var. integrifolia. It is apparently limited to Abies concolor forests above 1520 m (M. S. Baker 1953). Study of V. californica is needed to determine if taxonomic recognition is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
L. E. McKinney (1992) considered Viola affinis, and much of what botanists had called V. nephrophylla, to be essentially the same taxon. After studying additional specimens, reviewing literature (H. E. Ballard 1994; A. Haines 2001b), and discussions with others (J. Cayouette, H. E. Ballard, A. Haines, pers. comm.), he chose to maintain these as separate taxa. Reports of V. affinis in the Gulf coastal states based on specimens or photographs are usually attributable to V. missouriensis. Viola affinis reportedly hybridizes with V. hirsutula (= V. ×consobrina House), V. cucullata (= V. ×consocia House), V. brittoniana (= V. ×davisii House), V. sororia (= V. ×filicetorum Greene [as species]), V. sagittata var. sagittata (= V. ×hollickii House), and V. nephrophylla (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 120. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. californica, V. canadensis var. sitchensis, V. glabella var. remotifolia | V. affinis var. subarctica, V. crenulata, V. sororia subsp. affinis, V. sororia var. affinis, V. subviscosa, V. venustula |
Name authority | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 142. (1838) | Leconte: Ann. Lyceum Nat. Hist. New York 2: 138. (1826) |
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