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enable glossary links

pioneer violet, smooth yellow violet, stream or smooth yellow or yellow wood or wood or smooth yellow woodland violet, stream violet, violette glabre, yellow wood violet

Cascades early blue violet, common periwinkle, early blue violet, hook-spur violet, hookedspur violet, large periwinkle, sand violet, western dog violet, wild dog violet

Habit Plants perennial, caulescent, not stoloniferous, 3–38 cm. Plants perennial, caulescent, not stoloniferous, 1.8–30(–35) cm.
Stems

1–3, erect or prostrate, leafless proximally, leafy distally, glabrous or finely puberulent, on caudex from fleshy rhizome.

1–5, erect, ascending, or decumbent, sometimes later reclining to nearly prostrate, glabrous or puberulent, on caudex from subligneous rhizome.

Leaves

basal and cauline;

basal: 0–4(–7);

stipules ovate to obovate, margins entire, crenate, or serrate, usually glandular, apex acute to obtuse;

petiole 7–27.5 cm, glabrous or puberulent;

blade usually reniform to ovate, sometimes orbiculate, 3.3–8.5 × 2–9.3 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or finely puberulent;

cauline similar to basal except: usually restricted to distal ends of naked stems;

stipules ovate to oblong, margins erose or subserrate, often glandular, apex acute to acuminate;

petiole 0.2–2.9 cm, glabrous or finely puberulent;

blade ovate to deltate, 1.4–5.7 × 0.8–4.7 cm, base cordate to truncate, margins crenate to ± serrate, ciliate (sometimes limited to proximal half), apex acute.

basal and cauline;

basal: 1–4;

stipules linear to linear-lanceolate, margins entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 0.5–13.5 cm, glabrous or puberulent;

blade usually ovate or ovate-deltate to ovate-orbiculate, sometimes ± reniform or oblong, 0.5–6.9 × 0.4–5 cm, base cordate, subcordate, truncate, or attenuate, usually decurrent on petiole, margins crenate to crenulate or entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or sparsely to densely puberulent;

cauline similar to basal except: stipule margins lacerate to laciniate;

petiole 0.5–6.5 cm;

blade 0.6–5.5 × 0.4–4.7 cm.

Peduncles

2–8 cm, glabrous or pubescent.

1.7–13.8 cm, glabrous or puberulent.

Flowers

sepals linear-lanceolate, margins eciliate, auricles 0.5–1 mm;

petals deep lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 6–18 mm, spur yellow to greenish, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 0.5–2 mm;

petals light- to deep- to lavender-violet on both surfaces, rarely white, lower 3 usually white basally, dark violet-veined, lateral 2 (and sometimes upper 2) bearded, lowest 7–17(–23) mm, spur purple to violet or white, elongated, 5–7 mm, tip straight or pointed, curved up or lateral;

style head sparsely to densely bearded, sometimes beardless; cleistogamous flowers axillary.

Capsules

ovoid to ellipsoid, 7–13 mm, glabrous.

short-ovoid, 6–11 mm, glabrous.

Seeds

light to dark brown, shiny, 2–2.2 mm.

dark brown to olive-black, 1.5–2 mm.

2n

= 24.

Viola glabella

Viola adunca

Phenology Flowering Mar–Aug.
Habitat Damp, wet, or shady places in forests, stream banks
Elevation 0–2600 m (0–8500 ft)
Distribution
from FNA
AK; CA; ID; MT; OR; WA; AB; BC; Asia
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; IA; ID; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola glabella is similar in overall appearance to V. pubescens. The cauline leaves are sometimes described as appearing only near the apex of the stem (P. A. Munz 1959), but often a leaf occurs from a node on the stem below the apex.

Viola californica was described from collections made in Humboldt and Trinity counties, California, and may have arisen as a hybrid between V. glabella and V. lobata var. integrifolia. It is apparently limited to Abies concolor forests above 1520 m (M. S. Baker 1953). Study of V. californica is needed to determine if taxonomic recognition is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Viola adunca is polymorphic with over 50 named taxa. G. D. McPherson and J. G. Packer (1974) reported that diploid, triploid, and tetraploid races of V. adunca occur in Canada and northwestern United States. They found that diploid and tetraploid chromosome races can be distinguished morphologically based on style beards and on the size of guard cells and pollen grains and recommended taxonomic recognition of both races. In diploid races, the style projections are more or less cylindrical and about one-sixth the width of the style; in tetraploid races they are short-conical or globular, and about one-tenth or less than the width of the style beard.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants (4–)4.5–30(–35) cm; basal blades 1.3–6.9 × 1.2–5 cm; peduncles 3–13.8 cm; 0–3600 m; widespread in North America.
var. adunca
1. Plants 1.8–4.5(–6.5) cm; basal blades 0.5–1.7 × 0.4–1.4 cm; peduncles 1.7–5 cm; 2500–3800 m; restricted to Rocky Mountains (Colorado, Montana, Wyoming, British Columbia).
var. bellidifolia
Source FNA vol. 6, p. 131. FNA vol. 6, p. 119.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. adunca var. adunca, V. adunca var. bellidifolia
Synonyms V. californica, V. canadensis var. sitchensis, V. glabella var. remotifolia Lophion aduncum, V. canina var. adunca
Name authority Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 142. (1838) Smith: in A. Rees, Cycl. 37: Viola no. 63
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