Viola flettii |
Viola frank-smithii |
Viola missouriensis |
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Flett's violet, Olympic violet, rock violet |
Frank Smith violet, Frank-Smith's violet |
Missouri violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–15 cm. | Plants perennial, caulescent, not stoloniferous, 1.5–12.5 cm. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizomes thick, fleshy. |
Stems | 1–3, ascending to erect, mostly glabrous, on caudex from fleshy rhizome. |
1–4, decumbent or ascending to erect, glabrous, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 1–3; stipules linear-lanceolate, margins entire or with glandular processes, apex acuminate; petiole 1.5–9.7 cm, mostly glabrous; blade purple-tinted and –veined, broadly reniform to ovate, 0.9–2.4 × 1.2–4 cm, base cordate, margins finely crenate-serrate, eciliate, apex acute to obtuse, surfaces glabrous or sparsely pubescent along veins adaxially; cauline similar to basal except: stipules ovate to lanceolate, margins entire or shallowly laciniate; petiole 0.7–5.9 cm, usually glabrous; blade 0.8–2.1 × 1.2–3.1 cm. |
basal and cauline; basal: 2–3; stipules lanceolate, margins fimbriate, apex acute; petiole 1.5–6(–14) cm, glabrous; blade broadly ovate to deltate, 1.3–2.9(–4.1) × 1.3–2.9 cm, base cordate to truncate, margins crenate, eciliate, apex obtuse to rounded, surfaces glabrous; cauline similar to basal except: petiole 1.3–6.5 cm; blade 1.1–2.2 × 0.7–1.6 cm. |
basal, 1–8, ascending to erect; stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes distally glandular, apex acute; petiole 5–20 cm, glabrous; blade green abaxially, unlobed, usually narrowly to broadly deltate, 1.5–12 × 1.5–10 cm, not fleshy, base cordate or broadly cordate to truncate, margins ± crenate to serrate, ciliate or eciliate, apex acute to acuminate, surfaces glabrous, rarely pubescent. |
Peduncles | 1.8–7.1 cm, usually glabrous. |
2.5–7(–11.2) cm, glabrous. |
3–25 cm, usually glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals soft reddish violet on both surfaces, all with yellow area basally, lower 3 dark violet-veined, lateral 2 bearded, lowest with white around yellow area, 10–15 mm, spur yellow, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm; all petals usually pale purple, sometimes almost white adaxially, lateral 2 and lowest purple-veined, lateral and upper 2 violet abaxially, white basally, lateral 2 bearded, lowest 10–16(–18) mm, spur greenish to pale lime green, gibbous, 1.6–3 mm; style head bearded; cleistogamous flowers unknown. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest beardless, rarely lightly bearded, 15–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | ± spherical, 5–9 mm, glabrous. |
ovoid, 3–7 mm, glabrous. |
ellipsoid, 5–12 mm, glabrous. |
Seeds | dark brown to brownish purple, 2.5–3 mm. |
pale yellow, 2.4–2.8 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 54. |
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Viola flettii |
Viola frank-smithii |
Viola missouriensis |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. | Flowering Mar–May. |
Habitat | Alpine and subalpine rock crevices, vertical faces, talus slopes | Cracks, crevices, and narrow ledges of vertical limestone and dolomite rock faces, humid, shady places | Swamps, thickets, stream banks, alluvial woods |
Elevation | 1100–2000 m (3600–6600 ft) | 1600–2100 m (5200–6900 ft) | 50–2000 m (200–6600 ft) |
Distribution |
WA
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UT |
AL; AR; DC; DE; FL; GA; IA; IL; IN; KS; LA; MD; MN; MO; MS; NC; NE; NJ; NM; OK; SC; SD; TN; TX; VA; WI
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Discussion | Viola flettii is endemic to the Olympic Mountains of northwestern Washington. C. S. McCreary (2005) noted that although morphologically and ecologically distinct, V. cuneata, V. flettii, and V. ocellata are closely related. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola frank-smithii is endemic to Logan Canyon, Cache County, Utah. N. H. Holmgren (2005d) wrote that it likely has its ancestral ties with V. adunca. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
What many have treated as Viola affinis, especially in the southern Gulf coastal states, is likely to be V. missouriensis. N. H. Russell (1965) considered the two as likely part of a species complex inhabiting alluvial woods and wet areas and exhibiting the typical deltate leaf blade shape. L. E. McKinney (1992) considered V. missouriensis a variety of V. sororia. Viola missouriensis appears to have a closer affinity to V. affinis, as Russell suggested; current evidence suggests maintaining V. missouriensis as a separate species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 131. | FNA vol. 6, p. 138. |
Parent taxa | Violaceae > Viola | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | |||
Synonyms | V. candidula, V. langloisii, V. lucidifolia, V. sororia var. missouriensis | ||
Name authority | Piper: Erythea 6: 69. (1898) | N. H. Holmgren: Brittonia 44: 303, fig. 1E – H. (1992) | Greene: Pittonia 4: 141. (1900) |
Web links | |||