Viola flettii |
Viola cuneata |
Viola frank-smithii |
|
---|---|---|---|
Flett's violet, Olympic violet, rock violet |
northern two-eyed or wedge-leaf violet, northern two-eyed violet, wedge-leaf violet |
Frank Smith violet, Frank-Smith's violet |
|
Habit | Plants perennial, caulescent, not stoloniferous, 3–15 cm. | Plants perennial, caulescent, not stoloniferous, 2–25 cm. | Plants perennial, caulescent, not stoloniferous, 1.5–12.5 cm. |
Stems | 1–3, ascending to erect, mostly glabrous, on caudex from fleshy rhizome. |
1–3, usually erect, sometimes prostrate or ascending, glabrous, from shallow, fleshy rhizome or deep-seated caudex with fleshy roots. |
1–4, decumbent or ascending to erect, glabrous, on caudex from fleshy rhizome. |
Leaves | basal and cauline; basal: 1–3; stipules linear-lanceolate, margins entire or with glandular processes, apex acuminate; petiole 1.5–9.7 cm, mostly glabrous; blade purple-tinted and –veined, broadly reniform to ovate, 0.9–2.4 × 1.2–4 cm, base cordate, margins finely crenate-serrate, eciliate, apex acute to obtuse, surfaces glabrous or sparsely pubescent along veins adaxially; cauline similar to basal except: stipules ovate to lanceolate, margins entire or shallowly laciniate; petiole 0.7–5.9 cm, usually glabrous; blade 0.8–2.1 × 1.2–3.1 cm. |
basal and cauline; basal: 2–6; stipules lanceolate, margins entire or gland-toothed, apex acute; petiole 4.5–9.8 cm, glabrous; blade purple-veined, orbiculate-ovate to deltate, 1–4 × 2.7–3.8 cm, usually shiny, leathery, base cuneate, margins serrate, eciliate, apex acute, mucronulate, surfaces glabrous; cauline similar to basal except: stipules lanceolate to ovate, margins entire or erose to lacerate, with or without gland-tipped processes, apex acute to acuminate; petiole 0.5–20 cm; blade usually rhombic, sometimes orbiculate, 0.9–2.6 × 0.7–1.8 cm, margins crenate to ± serrate, apex acute to obtuse. |
basal and cauline; basal: 2–3; stipules lanceolate, margins fimbriate, apex acute; petiole 1.5–6(–14) cm, glabrous; blade broadly ovate to deltate, 1.3–2.9(–4.1) × 1.3–2.9 cm, base cordate to truncate, margins crenate, eciliate, apex obtuse to rounded, surfaces glabrous; cauline similar to basal except: petiole 1.3–6.5 cm; blade 1.1–2.2 × 0.7–1.6 cm. |
Peduncles | 1.8–7.1 cm, usually glabrous. |
1–10.5 cm, glabrous. |
2.5–7(–11.2) cm, glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals soft reddish violet on both surfaces, all with yellow area basally, lower 3 dark violet-veined, lateral 2 bearded, lowest with white around yellow area, 10–15 mm, spur yellow, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep reddish violet abaxially, white adaxially, upper 2 sometimes with purple patch basally, lateral 2 with purple patch basally beyond smaller yellow area, usually bearded, sometimes beardless, lowest purple-veined with yellow area basally, 8–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm; all petals usually pale purple, sometimes almost white adaxially, lateral 2 and lowest purple-veined, lateral and upper 2 violet abaxially, white basally, lateral 2 bearded, lowest 10–16(–18) mm, spur greenish to pale lime green, gibbous, 1.6–3 mm; style head bearded; cleistogamous flowers unknown. |
Capsules | ± spherical, 5–9 mm, glabrous. |
± spherical, 5–9 mm, glabrous. |
ovoid, 3–7 mm, glabrous. |
Seeds | dark brown to brownish purple, 2.5–3 mm. |
deep brown-purple, 2.1–3 mm. |
pale yellow, 2.4–2.8 mm. |
Viola flettii |
Viola cuneata |
Viola frank-smithii |
|
Phenology | Flowering Jun–Aug. | Flowering Mar–Sep. | Flowering May–Jul. |
Habitat | Alpine and subalpine rock crevices, vertical faces, talus slopes | Open pine or oak forests, often serpentine soil | Cracks, crevices, and narrow ledges of vertical limestone and dolomite rock faces, humid, shady places |
Elevation | 1100–2000 m (3600–6600 ft) | 600–2200 m (2000–7200 ft) | 1600–2100 m (5200–6900 ft) |
Distribution |
WA
|
CA; OR
|
UT |
Discussion | Viola flettii is endemic to the Olympic Mountains of northwestern Washington. C. S. McCreary (2005) noted that although morphologically and ecologically distinct, V. cuneata, V. flettii, and V. ocellata are closely related. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola cuneata often occurs on serpentine-derived soil and is closely related to V. flettii, V. hallii, and V. ocellata (V. B. Baird 1942). Originally described as beardless by Watson, the lateral petals of flowers in the same population can be prominently bearded or essentially beardless. The cauline blades of Viola cuneata are often vertical, especially early in season. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola frank-smithii is endemic to Logan Canyon, Cache County, Utah. N. H. Holmgren (2005d) wrote that it likely has its ancestral ties with V. adunca. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 129. | FNA vol. 6, p. 131. |
Parent taxa | Violaceae > Viola | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | |||
Name authority | Piper: Erythea 6: 69. (1898) | S. Watson: Proc. Amer. Acad. Arts 14: 290. (1879) | N. H. Holmgren: Brittonia 44: 303, fig. 1E – H. (1992) |
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