Viola flettii |
Viola bakeri |
Viola sororia |
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Flett's violet, Olympic violet, rock violet |
Baker's violet, yellow prairie violet |
common blue violet, common or downy or woolly blue violet, northern blue violet, northern violet, northern woodland violet, violette parente, woolly blue violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–15 cm. | Plants perennial, caulescent, not stoloniferous, 3–30 cm. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizomes thick, fleshy. |
Stems | 1–3, ascending to erect, mostly glabrous, on caudex from fleshy rhizome. |
1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–3; stipules linear-lanceolate, margins entire or with glandular processes, apex acuminate; petiole 1.5–9.7 cm, mostly glabrous; blade purple-tinted and –veined, broadly reniform to ovate, 0.9–2.4 × 1.2–4 cm, base cordate, margins finely crenate-serrate, eciliate, apex acute to obtuse, surfaces glabrous or sparsely pubescent along veins adaxially; cauline similar to basal except: stipules ovate to lanceolate, margins entire or shallowly laciniate; petiole 0.7–5.9 cm, usually glabrous; blade 0.8–2.1 × 1.2–3.1 cm. |
basal and cauline; basal: 1–4; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate; petiole 1–15.4 cm, glabrous or puberulent; blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins; cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections; petiole 1.5–7.5 cm; blade 1.9–6.7 × 0.5–1.6 cm. |
basal, 1–8, ascending to erect; stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes glandular distally, apex acute; petiole 2–25 cm, pubescent or glabrous; blade green abaxially, unlobed, ovate or broadly ovate to reniform, 2–5 × 2–10 cm, not fleshy, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually pubescent, rarely glabrous. |
Peduncles | 1.8–7.1 cm, usually glabrous. |
1.5–11.6 cm, glabrous or pubescent. |
3–25 cm, glabrous or sparsely pubescent. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals soft reddish violet on both surfaces, all with yellow area basally, lower 3 dark violet-veined, lateral 2 bearded, lowest with white around yellow area, 10–15 mm, spur yellow, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue- or dark purple-violet, reddish purple, or rarely white on both surfaces, usually white basally, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest bearded or beardless, 15–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | ± spherical, 5–9 mm, glabrous. |
spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent. |
ellipsoid, 5–12 mm, glabrous. |
Seeds | dark brown to brownish purple, 2.5–3 mm. |
light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 48. |
= 54. |
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Viola flettii |
Viola bakeri |
Viola sororia |
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Phenology | Flowering Jun–Aug. | Flowering May–Jul. | Flowering Mar–Jun. |
Habitat | Alpine and subalpine rock crevices, vertical faces, talus slopes | Wet and dry places in openings of coniferous forests | Dry to mesic habitats in woods, thickets, stream banks, moist prairies, pastures, disturbed ground, not in saturated soil |
Elevation | 1100–2000 m (3600–6600 ft) | 900–3800 m (3000–12500 ft) | 0–3000 m (0–9800 ft) |
Distribution |
WA
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CA; NV; OR; WA
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AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; BC; MB; NB; NF; NS; ON; PE; QC; SK
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Discussion | Viola flettii is endemic to the Olympic Mountains of northwestern Washington. C. S. McCreary (2005) noted that although morphologically and ecologically distinct, V. cuneata, V. flettii, and V. ocellata are closely related. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3). Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola sororia is similar to V. palmata in the high degree of phenotypic plasticity. The petal color is usually light to dark blue-violet. In V. sororia forma priceana (the Confederate violet) petals are grayish white with violet veins. Viola septentrionalis was recognized by N. L. Gil-Ad (1997) based on what he considered distinct capsule and seed morphology. His assumptions were based on one to four specimens. A. Haines (2011) alluded to the variability of V. sororia and indicated that a more northern form has often been called V. septentrionalis. He suggested that range-wide patterns of variation make it difficult or impractical to separate. We currently see no valid reason to recognize V. septentrionalis. Viola floridana was recognized by D. B. Ward (2006) as distinct from V. sororia based on being glabrous except for scattered hairs on petioles. Viola sororia reportedly hybridizes with V. cucullata (= V. ×bissellii House), V. hirsutula [= V. ×cordifolia (Nuttall) Schweinitz], V. pedatifida var. brittoniana (= V. ×insolita House), V. pedatifida var. pedatifida (= V. ×bernardii Greene), and V. sagittata var. sagittata (= V. ×conjugens Greene). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 121. | FNA vol. 6, p. 158. |
Parent taxa | Violaceae > Viola | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | |||
Synonyms | V. bakeri subsp. shastensis, V. nuttallii var. bakeri | V. chalcosperma, V. floridana, V. latiuscula, V. palmata var. sororia, V. papilionaceae, V. priceana, V. rosacea, V. septentrionalis, V. wilmattiae | |
Name authority | Piper: Erythea 6: 69. (1898) | Greene: Pittonia 3: 307. (1898) | Willdenow: Hort. Berol. 1: plate 72. (1806) |
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