Viola flettii |
Viola bakeri |
Viola canina |
|
---|---|---|---|
Flett's violet, Olympic violet, rock violet |
Baker's violet, yellow prairie violet |
dog violet, heath dog-violet |
|
Habit | Plants perennial, caulescent, not stoloniferous, 3–15 cm. | Plants perennial, caulescent, not stoloniferous, 3–30 cm. | Plants perennial, caulescent, not stoloniferous, 2–40 cm. |
Stems | 1–3, ascending to erect, mostly glabrous, on caudex from fleshy rhizome. |
1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome. |
1–5, ascending to erect, glabrous, on caudex from subligneous rhizome. |
Leaves | basal and cauline; basal: 1–3; stipules linear-lanceolate, margins entire or with glandular processes, apex acuminate; petiole 1.5–9.7 cm, mostly glabrous; blade purple-tinted and –veined, broadly reniform to ovate, 0.9–2.4 × 1.2–4 cm, base cordate, margins finely crenate-serrate, eciliate, apex acute to obtuse, surfaces glabrous or sparsely pubescent along veins adaxially; cauline similar to basal except: stipules ovate to lanceolate, margins entire or shallowly laciniate; petiole 0.7–5.9 cm, usually glabrous; blade 0.8–2.1 × 1.2–3.1 cm. |
basal and cauline; basal: 1–4; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate; petiole 1–15.4 cm, glabrous or puberulent; blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins; cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections; petiole 1.5–7.5 cm; blade 1.9–6.7 × 0.5–1.6 cm. |
cauline; stipules linear to lanceolate, margins ± lacerate to subentire, points often gland-tipped, apex acute; petiole 0.3–2.5 cm, glabrous; blade ovate to narrowly ovate, 1.2–5.2 × 1–2.7 cm, base cordate or deeply cordate to truncate, margins crenate, eciliate, apex acute to obtuse, abaxial surface glabrous or sparsely pubescent on veins, adaxial surface glabrous or sparsely pubescent. |
Peduncles | 1.8–7.1 cm, usually glabrous. |
1.5–11.6 cm, glabrous or pubescent. |
1–10 cm, glabrous or sparsely puberulent. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals soft reddish violet on both surfaces, all with yellow area basally, lower 3 dark violet-veined, lateral 2 bearded, lowest with white around yellow area, 10–15 mm, spur yellow, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 2–3 mm; petals usually blue- to gray-violet on both surfaces, rarely white, white basally, lower 3 dark violet-veined, lateral 2 bearded, lowest 15–25 mm, spur light yellow to light green or white, gibbous to usually elongated, 3–5 mm; style head beardless; cleistogamous flowers axillary. |
Capsules | ± spherical, 5–9 mm, glabrous. |
spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent. |
ovoid, 7.5–9 mm, glabrous. |
Seeds | dark brown to brownish purple, 2.5–3 mm. |
light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus. |
light to dark brown; 1.7–2.1 mm. |
2n | = 48. |
= 40. |
|
Viola flettii |
Viola bakeri |
Viola canina |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. | Flowering May–Jul. |
Habitat | Alpine and subalpine rock crevices, vertical faces, talus slopes | Wet and dry places in openings of coniferous forests | Sunny to shady, dry to slightly moist ground, heath lands, grazed or mown grassland, dunes, rock ledges, open woodland, disturbed areas (roadsides, railway banks, gravel pits), littoral vegetation of lakes and streams |
Elevation | 1100–2000 m (3600–6600 ft) | 900–3800 m (3000–12500 ft) | 20–300 m (100–1000 ft) |
Distribution |
WA
|
CA; NV; OR; WA
|
Greenland; Europe; Asia |
Discussion | Viola flettii is endemic to the Olympic Mountains of northwestern Washington. C. S. McCreary (2005) noted that although morphologically and ecologically distinct, V. cuneata, V. flettii, and V. ocellata are closely related. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3). Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola canina was possibly introduced in Greenland (T. Marcussen and T. Karlsson et al. 2010). Hybrids of Viola canina are known with the European species V. mirabilis Linnaeus, V. pumila Chaix, V. reichenbachiana Jordan ex Boreau, V. riviniana Reichenbach, V. rupestris F. W. Schmidt (both subspecies), V. stagnina Kitaibel, and V. uliginosa Besser. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 131. | FNA vol. 6, p. 121. | FNA vol. 6, p. 127. |
Parent taxa | Violaceae > Viola | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | |||
Synonyms | V. bakeri subsp. shastensis, V. nuttallii var. bakeri | V. canina var. montana, V. montana, V. nemoralis | |
Name authority | Piper: Erythea 6: 69. (1898) | Greene: Pittonia 3: 307. (1898) | Linnaeus: Sp. Pl. 2: 935. (1753) |
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