Viola douglasii |
Viola walteri |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
prostrate blue violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, caulescent, stoloniferous, 5–19 cm; stolons green or reddish, leafy, sometimes rooting at nodes, becoming lignified in age. | ||||
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
1–5, prostrate, spreading, finely puberulent, from current and/or previous year’s growth, on usually vertical, fleshy rhizome, rooting and forming rosettes at or near tip; rooted rosettes often develop into an erect, fleshy caudex from which new stems are produced. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal and cauline; basal: 3–6; stipules lanceolate, margins laciniate, projections often long-filamentous, apex long-acuminate; petiole 2.3–7.3 cm, glabrous or pubescent; blade often purple-spotted abaxially and/or adaxially, ovate to reniform, 1.2–5 × 1.6–3.6 cm, base deeply to broadly cordate, margins crenulate to serrate, ciliate or eciliate, apex rounded to obtuse, surfaces glabrous or pubescent; cauline similar to basal except: stipules ovate to lanceolate, margins laciniate; petiole 1–3.5 cm; blade 1.3–2.9 × 1.4–3.2 cm. |
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Peduncles | 2–12.5 cm, glabrous or puberulent. |
5–9.6 cm, glabrous or pubescent. |
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Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins mostly eciliate, auricles 0.5–1 mm; petals pale to bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 and often upper 2 and lowest bearded, lowest 15–18 mm, spur white, gibbous to usually elongated, 3–5 mm; style head bearded; cleistogamous flowers axillary. |
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Capsules | spherical to oblong, 5–12 mm, glabrous. |
ovoid to ellipsoid, 5–7 mm, glabrous. |
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Seeds | light brown, 2.8–3.3 mm. |
brown, 1–1.5 mm. |
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2n | = 24, 48. |
= 20. |
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Viola douglasii |
Viola walteri |
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Phenology | Flowering Feb–Jul. | |||||
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | |||||
Elevation | 20–2300 m (100–7500 ft) | |||||
Distribution |
CA; OR; Mexico (Baja California)
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AL; AR; FL; GA; KY; LA; MD; MS; NC; OH; PA; SC; TN; TX; VA; WV
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 163. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. chrysantha | |||||
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | House: Torreya 6: 172. (1906) | ||||
Web links |