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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

prostrate blue violet

Habit Plants perennial, caulescent, not stoloniferous, 3–20 cm. Plants perennial, caulescent, stoloniferous, 5–19 cm; stolons green or reddish, leafy, sometimes rooting at nodes, becoming lignified in age.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

1–5, prostrate, spreading, finely puberulent, from current and/or previous year’s growth, on usually vertical, fleshy rhizome, rooting and forming rosettes at or near tip; rooted rosettes often develop into an erect, fleshy caudex from which new stems are produced.

Leaves

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

basal and cauline;

basal: 3–6;

stipules lanceolate, margins laciniate, projections often long-filamentous, apex long-acuminate;

petiole 2.3–7.3 cm, glabrous or pubescent;

blade often purple-spotted abaxially and/or adaxially, ovate to reniform, 1.2–5 × 1.6–3.6 cm, base deeply to broadly cordate, margins crenulate to serrate, ciliate or eciliate, apex rounded to obtuse, surfaces glabrous or pubescent;

cauline similar to basal except: stipules ovate to lanceolate, margins laciniate;

petiole 1–3.5 cm;

blade 1.3–2.9 × 1.4–3.2 cm.

Peduncles

2–12.5 cm, glabrous or puberulent.

5–9.6 cm, glabrous or pubescent.

Flowers

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate to ovate, margins mostly eciliate, auricles 0.5–1 mm;

petals pale to bluish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 and often upper 2 and lowest bearded, lowest 15–18 mm, spur white, gibbous to usually elongated, 3–5 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

spherical to oblong, 5–12 mm, glabrous.

ovoid to ellipsoid, 5–7 mm, glabrous.

Seeds

light brown, 2.8–3.3 mm.

brown, 1–1.5 mm.

2n

= 24, 48.

= 20.

Viola douglasii

Viola walteri

Phenology Flowering Feb–Jul.
Habitat Vernally moist grassy slopes and flats, often serpentine soil (except Oregon)
Elevation 20–2300 m (100–7500 ft)
Distribution
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; KY; LA; MD; MS; NC; OH; PA; SC; TN; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades pubescent, hairs scattered throughout or concentrated along veins, adaxial surfaces, or toward margins; stipule margin processes 1/2+ stipule length.
var. walteri
1. Leaf blades glabrous or pubescent, hairs usually confined to adaxial surfaces; stipule margin processes to 1/2 stipule length.
var. appalachiensis
Source FNA vol. 6, p. 129. FNA vol. 6, p. 163.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa
Subordinate taxa
V. walteri var. appalachiensis, V. walteri var. walteri
Synonyms V. chrysantha
Name authority Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) House: Torreya 6: 172. (1906)
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