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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

Carolina violet, southern woolly violet

Habit Plants perennial, caulescent, not stoloniferous, 3–20 cm. Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome thick, fleshy.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

Leaves

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

basal, 4–9, prostrate to ascending;

stipules linear-lanceolate, margins entire, apex acute;

petiole 3–10 cm, densely pubescent;

blade unlobed, reniform or ovate to elliptic, 1–8 × 1–5.5 cm, base cordate, margins serrate, ciliate, apex rounded to acute, mucronulate, surfaces densely pubescent.

Peduncles

2–12.5 cm, glabrous or puberulent.

4–10 cm, puberulent.

Flowers

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate to ovate, margins ciliate, auricles 1–2 mm;

petals light to dark blue-violet on both surfaces, lower 3 white basally and dark violet-veined, lateral 2 bearded, spur sometimes bearded, lowest 10–20 mm, spur usually white, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on ascending to erect peduncles.

Capsules

spherical to oblong, 5–12 mm, glabrous.

ellipsoid, 6–10 mm, glabrous.

Seeds

light brown, 2.8–3.3 mm.

beige, mottled to bronze, or dark brown, 1.5–2 mm.

2n

= 24, 48.

= 54.

Viola douglasii

Viola villosa

Phenology Flowering Feb–Jul. Flowering Apr–Jun.
Habitat Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) Sandy, pine-oak and pine-oak-hickory woods and disturbed ground
Elevation 20–2300 m (100–7500 ft) 10–300 m (0–1000 ft)
Distribution
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Much of the foliage of Viola villosa remains green throughout the winter (V. B. Baird 1942).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 129. FNA vol. 6, p. 163.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. walteri
Synonyms V. chrysantha V. alabamensis
Name authority Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) Walter: Fl. Carol., 219. (1788)
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