Viola douglasii |
Viola subsinuata |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
early blue violet, wavy-leaf violet, wood violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, acaulescent, not stoloniferous, 10–30 cm; rhizome thick, fleshy. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal, 2–11, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 5–25 cm, glabrous or pubescent; mid-season leaf blades incised or lobed throughout, earliest leaf blades lobed (plants homophyllous), similar to mid-season blades, blade 5–9(–16)-lobed, sinuses usually narrower, shallower toward leaf base, middle and lateral blade lobes differ in width and/or shape, middle lobes narrowly deltate to narrowly elliptic, lateral lobes narrowly elliptic, lanceolate, or falcate, 1–11 × 1–12 cm, base truncate to cordate, margins entire or crenate, ciliate or eciliate, apex acute, mucronulate, surfaces usually pubescent throughout or on veins. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
5–15 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet on both surfaces, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, lowest sometimes bearded, 15–25 mm, spur color same as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
ellipsoid, 8–12 mm, glabrous. |
Seeds | light brown, 2.8–3.3 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 24, 48. |
= 54. |
Viola douglasii |
Viola subsinuata |
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Phenology | Flowering Feb–Jul. | Flowering Apr–Jun. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Rich woods |
Elevation | 20–2300 m (100–7500 ft) | 100–3000 m (300–9800 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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CT; DC; DE; GA; IL; KY; MA; MD; MI; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; ON
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
L. E. McKinney (1992) described the misconceptions surrounding Viola subsinuata that many came to think of as the heterophyllous V. palmata, whereas V. subsinuata is homophyllous. As E. Brainerd (1910, 1921) pointed out, heterophylly versus homophylly is an important and steadfast character difference in acaulescent Viola species. N. L. Gil-Ad (1997) chose not to recognize V. subsinuata, suggesting that it represented a hybrid or introgressant. H. E. Ballard (2000) and A. Haines (2011) both recognized V. subsinuata as a species. Viola subsinuata is a state historical species in Rhode Island, where it was last documented in 1941 (R. W. Enser, http://rinhs.org/wp-content/uploads/ri_rare_plants_2007.pdf). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 159. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | V. emarginata var. subsinuata, V. palmata var. angelliae |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | (Greene) Greene: Pittonia 4: 4. (1899) |
Web links |