Viola douglasii |
Viola sororia |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
common blue violet, common or downy or woolly blue violet, northern blue violet, northern violet, northern woodland violet, violette parente, woolly blue violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizomes thick, fleshy. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal, 1–8, ascending to erect; stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes glandular distally, apex acute; petiole 2–25 cm, pubescent or glabrous; blade green abaxially, unlobed, ovate or broadly ovate to reniform, 2–5 × 2–10 cm, not fleshy, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually pubescent, rarely glabrous. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
3–25 cm, glabrous or sparsely pubescent. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue- or dark purple-violet, reddish purple, or rarely white on both surfaces, usually white basally, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest bearded or beardless, 15–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
ellipsoid, 5–12 mm, glabrous. |
Seeds | light brown, 2.8–3.3 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 24, 48. |
= 54. |
Viola douglasii |
Viola sororia |
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Phenology | Flowering Feb–Jul. | Flowering Mar–Jun. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Dry to mesic habitats in woods, thickets, stream banks, moist prairies, pastures, disturbed ground, not in saturated soil |
Elevation | 20–2300 m (100–7500 ft) | 0–3000 m (0–9800 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WI; WV; BC; MB; NB; NF; NS; ON; PE; QC; SK
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola sororia is similar to V. palmata in the high degree of phenotypic plasticity. The petal color is usually light to dark blue-violet. In V. sororia forma priceana (the Confederate violet) petals are grayish white with violet veins. Viola septentrionalis was recognized by N. L. Gil-Ad (1997) based on what he considered distinct capsule and seed morphology. His assumptions were based on one to four specimens. A. Haines (2011) alluded to the variability of V. sororia and indicated that a more northern form has often been called V. septentrionalis. He suggested that range-wide patterns of variation make it difficult or impractical to separate. We currently see no valid reason to recognize V. septentrionalis. Viola floridana was recognized by D. B. Ward (2006) as distinct from V. sororia based on being glabrous except for scattered hairs on petioles. Viola sororia reportedly hybridizes with V. cucullata (= V. ×bissellii House), V. hirsutula [= V. ×cordifolia (Nuttall) Schweinitz], V. pedatifida var. brittoniana (= V. ×insolita House), V. pedatifida var. pedatifida (= V. ×bernardii Greene), and V. sagittata var. sagittata (= V. ×conjugens Greene). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 158. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | V. chalcosperma, V. floridana, V. latiuscula, V. palmata var. sororia, V. papilionaceae, V. priceana, V. rosacea, V. septentrionalis, V. wilmattiae |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Willdenow: Hort. Berol. 1: plate 72. (1806) |
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