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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

common dog-violet, dog violet, wood violet

Habit Plants perennial, caulescent, not stoloniferous, 3–20 cm. Plants perennial, caulescent, not stoloniferous, 1.8–30 cm.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

1–5, erect, ascending, or decumbent, glabrous or puberulent, on caudex from subligneous rhizome.

Leaves

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

basal and cauline;

basal: 3–5;

stipules subulate or lanceolate, margins fimbriate, apex acute to acuminate;

petiole 2.5–12 cm, usually glabrous, rarely puberulent;

blade ovate to reniform, 2.3–5.5 × 2.6–5.3 cm, base cordate to deeply cordate, margins crenate, eciliate, apex acute or obtuse, abaxial surface glabrous or sparsely puberulent, adaxial surface usually sparsely puberulent, sometimes glabrous;

cauline similar to basal except: stipules ovate or lanceolate, margins fimbriate, apex long-acuminate;

petiole 1.3–6 cm;

blade ovate or broadly ovate, 1.8–5 × 1.5–4.2 cm, apex acute.

Peduncles

2–12.5 cm, glabrous or puberulent.

2.8–10 cm, glabrous, sometimes puberulent.

Flowers

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate, margins eciliate;

auricles 1.8–2.5 mm (enlarged in fruit);

petals violet on both surfaces, lower 3 usually white basally, rarely violet, dark violet-veined, lateral 2 bearded, lowest 13–17 mm, spur white, rarely violet, elongated, 5–7 mm, tip straight or hooked, blunt;

style head beardless (surface papillose); cleistogamous flowers axillary.

Capsules

spherical to oblong, 5–12 mm, glabrous.

ovoid or oblong, 8–12 mm, glabrous.

Seeds

light brown, 2.8–3.3 mm.

pale to light brown, 1.8–2.1 mm.

2n

= 24, 48.

= 40.

Viola douglasii

Viola riviniana

Phenology Flowering Feb–Jul. Flowering Mar–Sep.
Habitat Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) Roadsides and trails, sidewalk cracks, parks
Elevation 20–2300 m (100–7500 ft) 0–200 m (0–700 ft)
Distribution
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC; Eurasia; nw Africa [Introduced in North America; introduced in Australia]
Discussion

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola riviniana has become established in several states on the Pacific Coast. It is cultivated and sold through nurseries in the United States. In the nursery trade in California and elsewhere, it is incorrectly referred to incorrectly as V. labradorica ‘Purpurea.’ In some plants shoots arise from adventitious buds on the roots (A. R. Clapham et al. 1987; T. Marcussen and T. Karlsson 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 129. FNA vol. 6, p. 154.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. chrysantha
Name authority Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) Reichenbach: Iconogr. Bot. Pl. Crit. 1: 81, plate 95. (1823)
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