Viola douglasii |
Viola ocellata |
|
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
pinto violet, two-eyed violet, western heart's ease, western heart's ease or two-eyed or southern two-eyed violet |
|
Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, caulescent, not stoloniferous, 2–37 cm. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
1–3, ascending to erect, ± glabrous or usually puberulent, on caudex from shallow, fleshy rhizome or deep-seated caudex with fleshy roots. |
Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal and cauline; basal: 1–6; stipules deltate, margins laciniate, apex usually long-acuminate; petiole 3.7–10 cm, puberulent; blade ovate, deltate, or subreniform, 1–6 × 1.2–4 cm, base usually cordate to subcordate, margins crenate, usually ciliate, apex acute, mucronulate, surfaces usually sparsely puberulent, glabrous early; cauline similar to basal except: stipules lanceolate, margins ±fimbriate, sometimes stipitate-glandular, sometimes entire, erose, or ± laciniate; petiole 0.4–9 cm; blade ovate to deltate, 1.6–4.4 × 1.1–3.6 cm, base subcordate to truncate, margins crenate to ± serrate, ciliate, apex acute to obtuse. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
1–10 cm, puberulent. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals white adaxially, upper 2 and sometimes lower 3 deep reddish violet abaxially, all with yellow area basally, lateral 2 with purple patch basally distal to smaller yellow area, bearded, lowest with yellow patch basally, purple-veined, 5–15 mm, spur yellow or greenish, gibbous, 1–2.5 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
spherical to spherical-ovoid, 5–11 mm, minutely scabrous. |
Seeds | light brown, 2.8–3.3 mm. |
brown-purple, ca. 2 mm. |
2n | = 24, 48. |
= 12. |
Viola douglasii |
Viola ocellata |
|
Phenology | Flowering Feb–Jul. | Flowering Mar–Jul. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Rocky areas, grassy banks, thickets, often on serpentine soil |
Elevation | 20–2300 m (100–7500 ft) | 100–1100 m (300–3600 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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CA; OR
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 140. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Torrey & A. Gray: Fl. N. Amer. 1: 142. (1838) |
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