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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

pinto violet, two-eyed violet, western heart's ease, western heart's ease or two-eyed or southern two-eyed violet

Habit Plants perennial, caulescent, not stoloniferous, 3–20 cm. Plants perennial, caulescent, not stoloniferous, 2–37 cm.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

1–3, ascending to erect, ± glabrous or usually puberulent, on caudex from shallow, fleshy rhizome or deep-seated caudex with fleshy roots.

Leaves

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

basal and cauline;

basal: 1–6;

stipules deltate, margins laciniate, apex usually long-acuminate;

petiole 3.7–10 cm, puberulent;

blade ovate, deltate, or subreniform, 1–6 × 1.2–4 cm, base usually cordate to subcordate, margins crenate, usually ciliate, apex acute, mucronulate, surfaces usually sparsely puberulent, glabrous early;

cauline similar to basal except: stipules lanceolate, margins ±fimbriate, sometimes stipitate-glandular, sometimes entire, erose, or ± laciniate;

petiole 0.4–9 cm;

blade ovate to deltate, 1.6–4.4 × 1.1–3.6 cm, base subcordate to truncate, margins crenate to ± serrate, ciliate, apex acute to obtuse.

Peduncles

2–12.5 cm, glabrous or puberulent.

1–10 cm, puberulent.

Flowers

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals white adaxially, upper 2 and sometimes lower 3 deep reddish violet abaxially, all with yellow area basally, lateral 2 with purple patch basally distal to smaller yellow area, bearded, lowest with yellow patch basally, purple-veined, 5–15 mm, spur yellow or greenish, gibbous, 1–2.5 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

spherical to oblong, 5–12 mm, glabrous.

spherical to spherical-ovoid, 5–11 mm, minutely scabrous.

Seeds

light brown, 2.8–3.3 mm.

brown-purple, ca. 2 mm.

2n

= 24, 48.

= 12.

Viola douglasii

Viola ocellata

Phenology Flowering Feb–Jul. Flowering Mar–Jul.
Habitat Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) Rocky areas, grassy banks, thickets, often on serpentine soil
Elevation 20–2300 m (100–7500 ft) 100–1100 m (300–3600 ft)
Distribution
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 129. FNA vol. 6, p. 140.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. chrysantha
Name authority Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) Torrey & A. Gray: Fl. N. Amer. 1: 142. (1838)
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