Viola douglasii |
Viola nuttallii |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
Nuttall's or Nuttall's yellow or yellow prairie violet, Nuttall's violet, violette de Nuttall, yellow prairie violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, caulescent, not stoloniferous, 2–27 cm. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
1–6, ascending to erect, leafy proximally and distally, ca. 1/2 subterranean, puberulent, on caudex from usually vertical, subligneous rhizome. |
Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal and cauline; basal: 1–6; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute, few-toothed or lobed; petiole 2–17 cm, glabrous or minutely puberulent; blade lanceolate, ovate, or elliptic, 1–9 × 0.6–2.5 cm, base attenuate, margins entire or serrulate, sometimes sinuate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules adnate to or free from petiole, linear to linear-lanceolate or linear-oblong, sometimes leaflike, margins usually entire, rarely laciniate or glandular, apex acute to acuminate; petiole 2–7 cm; blade 1.4–7.2(–10) × 1.1–2.3 cm, length 1.3–4.4 times width, apex acute. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
3–13 cm, glabrous or puberulent. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals linear-lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially or on both surfaces, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–13 mm, spur yellow, gibbous, 0.5–1.5 mm; style head bearded; cleistogamous flowers axillary. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
subglobose to ovoid, 4–10 mm, usually glabrous, rarely finely puberulent. |
Seeds | light brown, 2.8–3.3 mm. |
medium brown, 2–3.2 mm, elaiosome extending over 1/3 length of seed and completely covering funiculus. |
2n | = 24, 48. |
= 24. |
Viola douglasii |
Viola nuttallii |
|
Phenology | Flowering Feb–Jul. | Flowering Apr–Jun. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Sagebrush flats, prairie grasslands, dry stream banks, juniper woodlands, scree slopes |
Elevation | 20–2300 m (100–7500 ft) | 400–2600 m (1300–8500 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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AZ; CO; ID; KS; MN; MT; ND; NE; NM; SD; UT; WY; AB; MB; SK
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
D. M. Fabijan et al. (1987) stated that Viola nuttallii showed no evidence of hybridization even when growing with V. vallicola east of the Rocky Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 140. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | Crocion nuttallii |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Pursh: Fl. Amer. Sept. 1: 174. (1813) |
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