Viola douglasii |
Viola lobata |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
moose horn violet, pine violet, yellow wood violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, caulescent, not stoloniferous, 5–46 cm. | ||||
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
1–3, erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal and cauline; basal: 0–2, unlobed or palmately 3–11-lobed; stipules ovate to lanceolate, margins ± entire or serrate, apex acute; petiole 5–27 cm, glabrous or puberulent; blade sometimes glaucous, deltate to reniform, 3.5–8.5 × 4.5–13.5 cm, base cordate, truncate, or attenuate, margins on unlobed leaves coarsely dentate-serrate, margins on lobed leaves usually entire, sometimes few-toothed, ciliate (sometimes only proximal 1/2) or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: distal on naked stems, unlobed and divided leaves can occur on same plant, if divided, palmately 3–12-lobed; stipules sometimes large and ± leaflike, margins entire, lacerate, or laciniate, sometimes with gland-tipped projections, apex acute to acuminate; petiole 0.2–8.8 cm; blade reniform to reniform-cordate, ± ovate, deltate, or rhombic, 1.5–5.5 × 1.4–10 cm, base cordate, subcordate, truncate, or attenuate, margins entire, crenate-serrate, or dentate, or coarsely lacerate to deeply serrate, often entire distally, ciliate or rarely eciliate, apex acute to obtuse, often long-tapered, mucronulate. |
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Peduncles | 2–12.5 cm, glabrous or puberulent. |
2–13 cm, glabrous or pubescent. |
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Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins ciliate or eciliate, auricles 0.1–1 mm; petals deep lemon-yellow adaxially, usually upper 2 and sometimes lateral 2 brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined basally, lateral 2 bearded, lowest 8–19 mm, spur yellow to greenish, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers axillary. |
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Capsules | spherical to oblong, 5–12 mm, glabrous. |
ellipsoid-ovoid, 6–16 mm, glabrous. |
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Seeds | light brown, 2.8–3.3 mm. |
light brown, blotched or streaked with brown, shiny, 2.1–2.7 mm. |
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2n | = 24, 48. |
= 12. |
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Viola douglasii |
Viola lobata |
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Phenology | Flowering Feb–Jul. | |||||
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | |||||
Elevation | 20–2300 m (100–7500 ft) | |||||
Distribution |
CA; OR; Mexico (Baja California)
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CA; OR; nw Mexico
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). V. B. Baird (1942) noted that Viola lobata was more closely related to V. tripartita than to any western Viola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 137. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. chrysantha | |||||
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Bentham: Pl. Hartw., 298. (1849) | ||||
Web links |