Viola douglasii
Viola guadalupensis
Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet
Guadalupe Mountains violet, Guadalupe violet
1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.
1–5, decumbent to erect, leafy proximally and distally, glabrous, on caudex from fleshy rhizome.
basal and cauline;
basal: 1–6, bipinnately compound, leaflets 3–5;
stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;
petiole 5–6.8 cm, glabrous or puberulent;
blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;
cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;
petiole 0.9–4 cm;
blade 1.1–4.1 × 1–3.6 cm.
cauline;
stipules lanceolate to ovate- or oblong-lanceolate or linear, margins sparingly glandular-fimbriate, apex acute;
petiole 2–6 cm, glabrous;
blade ovate to ovate-deltate or ovate-lanceolate, 1.2–2.4 × 0.7–1.3 cm, base broadly cuneate to rounded or truncate, margins entire or with 1–3 crenations on proximal 1/2, eciliate, apex acute to rounded, surfaces glabrous, sometimes with a few short hairs on veins abaxially.
2–12.5 cm, glabrous or puberulent.
3.5–6 cm, glabrous.
sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;
petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;
style head bearded; cleistogamous flowers absent.
sepals linear to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm;
petals deep lemon-yellow adaxially, upper 2 reddish brown abaxially, lateral 2 and lowest dark brown-veined basally, lateral 2 bearded, lowest 7–11 mm, spur yellow, gibbous, 1–1.4 mm;
style head bearded; cleistogamous flowers absent.
spherical to oblong, 5–12 mm, glabrous.
ovoid, 3–4.5 mm, glabrous.
light brown, 2.8–3.3 mm.
light brown, ± 2 mm.
= 24, 48.
= 24.
Viola douglasii
Viola guadalupensis
Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Of conservation concern.
Viola guadalupensis is known only from the eastern rim of the Guadalupe Mountains in Culberson County. Powell and Wauer noted that it is the only yellow-flowered violet known in the Guadalupe Mountains and appears to be related to V. nuttallii and V. vallicola. K. W. Allred (2008) stated that a report of this species in New Mexico by J. T. Kartesz and C. A. Meacham (1999) requires verification. K. Haskins (pers. comm.) reported that experiments are currently being conducted to propagate plants of V. guadalupensis via cell tissue culture.
Chloroplast (trnL-F spacer) and low-copy nuclear gene (GPI) phylogenies indicate that Viola guadalupensis is an alloploid that originated through hybridization between an unidentified member of subsect. Canadenses (the paternal parent) and a member of the V. nuttallii complex
(the maternal parent), of sect. Chamaemelanium (T. Marcussen et al. 2011). Evidence reported by these authors from a fossil-calibrated relaxed clock dating analysis showed the estimated maximum age of V. guadalupensis to be (5.7–)8.6(–11.6) million years.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
CA
OR
Flora NW
PNW Herbaria
WildflowerSearch
iNaturalist
USDA Plants Database
