Viola douglasii |
Viola frank-smithii |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
Frank Smith violet, Frank-Smith's violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, caulescent, not stoloniferous, 1.5–12.5 cm. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
1–4, decumbent or ascending to erect, glabrous, on caudex from fleshy rhizome. |
Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal and cauline; basal: 2–3; stipules lanceolate, margins fimbriate, apex acute; petiole 1.5–6(–14) cm, glabrous; blade broadly ovate to deltate, 1.3–2.9(–4.1) × 1.3–2.9 cm, base cordate to truncate, margins crenate, eciliate, apex obtuse to rounded, surfaces glabrous; cauline similar to basal except: petiole 1.3–6.5 cm; blade 1.1–2.2 × 0.7–1.6 cm. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
2.5–7(–11.2) cm, glabrous. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm; all petals usually pale purple, sometimes almost white adaxially, lateral 2 and lowest purple-veined, lateral and upper 2 violet abaxially, white basally, lateral 2 bearded, lowest 10–16(–18) mm, spur greenish to pale lime green, gibbous, 1.6–3 mm; style head bearded; cleistogamous flowers unknown. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
ovoid, 3–7 mm, glabrous. |
Seeds | light brown, 2.8–3.3 mm. |
pale yellow, 2.4–2.8 mm. |
2n | = 24, 48. |
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Viola douglasii |
Viola frank-smithii |
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Phenology | Flowering Feb–Jul. | Flowering May–Jul. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Cracks, crevices, and narrow ledges of vertical limestone and dolomite rock faces, humid, shady places |
Elevation | 20–2300 m (100–7500 ft) | 1600–2100 m (5200–6900 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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UT |
Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola frank-smithii is endemic to Logan Canyon, Cache County, Utah. N. H. Holmgren (2005d) wrote that it likely has its ancestral ties with V. adunca. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 131. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | N. H. Holmgren: Brittonia 44: 303, fig. 1E – H. (1992) |
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