Viola douglasii |
Viola egglestonii |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
cedar glade or glade violet, glade violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, acaulescent, not stoloniferous, 5–20 cm; rhizome thick, fleshy. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal, 3, prostrate to ascending; stipules linear-lanceolate, margins entire, apex acute; petiole 1.5–7 cm, usually glabrous; earliest leaf blades ± deltate or 3-lobed, mid-season blades 5–9-lobed, 1–9 × 1–10 cm, base truncate to cuneate, middle lobe lanceolate or spatulate to narrowly ovate, lateral lobes lanceolate or spatulate to falcate, margins serrate, sometimes with deltate or falcate appendages or teeth, ciliate, apex acute, surfaces usually glabrous, rarely pubescent. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
2–15 cm, usually glabrous, rarely pubescent. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 0.5–1 mm; petals light to dark blue-violet on both surfaces, lower 3 and sometimes upper 2 white basally, lower 3 darker violet-veined, lateral 2 densely bearded, lowest 10–15 mm, spur usually lilac, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate peduncles. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
ellipsoid, 11–14 mm, glabrous. |
Seeds | light brown, 2.8–3.3 mm. |
beige, mottled to bronze, 2–3 mm. |
2n | = 24, 48. |
= 54. |
Viola douglasii |
Viola egglestonii |
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Phenology | Flowering Feb–Jul. | Flowering Mar–May. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Limestone glades and barrens |
Elevation | 20–2300 m (100–7500 ft) | 100–200 m (300–700 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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AL; GA; IN; KY; TN |
Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
N. H. Russell (1965) was the first to consider Viola egglestonii similar to V. septemloba. After analyzing leaf-blade lobing patterns, L. E. McKinney considered it a subspecies of V. septemloba. Although similarities exist between these taxa, we maintain them as distinct species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 129. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | V. septemloba subsp. egglestonii |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Brainerd: Bull. Torrey Bot. Club 37: 526, plates 34, 35. (1910) |
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