Viola douglasii |
Viola canadensis |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
Canada violet, Canada white violet, Canadian violet, Canadian white violet, rugose violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, caulescent, not stoloniferous, 3–46(–60) cm, with branching rhizomes forming colonies or not. | ||||||||
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
1–3(4), usually erect to ascending, glabrous or puberulent, from fleshy or subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal and cauline; basal: 1–5; stipules ± oblong, ovate, or lanceolate, margins entire, sometimes glandular, apex acuminate to cuspidate; petiole 1.1–23 cm, glabrous or puberulent; blade ovate to broadly ovate or ovate-reniform, 0.7–12.4 × 0.9–11.1(–12.3) cm, base cordate, subcordate, or truncate, margins crenate, crenulate, or serrulate, ciliate (sometimes only on proximal 1/2) or eciliate, apex acute to acuminate, surfaces glabrous or puberulent (often only on veins); cauline similar to basal except: stipules also deltate, margins also erose or laciniate, apex acute, long-acuminate to cuspidate, or ± truncate, occasionally 2- or 3-fid; petiole 0.1–6.9(–15.2) cm; blade ovate to deltate, 1.2–7.7 × 0.8–7.8 cm, base cordate to truncate, margins crenate or crenulate to ± serrulate. |
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Peduncles | 2–12.5 cm, glabrous or puberulent. |
1–6.1 cm, glabrate to puberulent, sometimes glabrous below bracteoles. |
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Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins usually eciliate, auricles 0.5–1.3 mm; petals white adaxially, upper 2 and lower 3 tinged soft reddish violet abaxially, rarely white on both surfaces, all petals usually with yellow patch basally, lower 3 usually purple-veined, lateral 2 bearded, lowest 5.5–20 mm, spur white, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary or absent. |
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Capsules | spherical to oblong, 5–12 mm, glabrous. |
ovoid to ellipsoid, 2.5–10 mm, sometimes muriculate, glabrous or puberulent. |
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Seeds | light brown, 2.8–3.3 mm. |
brown to dark brown or purplish black, 1.5–2.5 mm. |
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2n | = 24, 48. |
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Viola douglasii |
Viola canadensis |
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Phenology | Flowering Feb–Jul. | |||||||||
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | |||||||||
Elevation | 20–2300 m (100–7500 ft) | |||||||||
Distribution |
CA; OR; Mexico (Baja California)
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AK; AL; AR; AZ; CO; CT; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OR; PA; RI; SC; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; NT; ON; QC; SK; YT
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 125. | ||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | V. chrysantha | |||||||||
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Linnaeus: Sp. Pl. 2: 936. (1753) | ||||||||
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