Viola douglasii |
Viola blanda |
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Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet |
sweet white violet, violette agréable |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–20 cm. | Plants perennial, acaulescent, stoloniferous, 3–20 cm; stolons pale, often rooting and leafy at nodes; rhizome short, slender, fleshy. |
Stems | 1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–6, bipinnately compound, leaflets 3–5; stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate; petiole 5–6.8 cm, glabrous or puberulent; blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent; cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute; petiole 0.9–4 cm; blade 1.1–4.1 × 1–3.6 cm. |
basal, 2–9, prostrate to ascending; stipules linear-lanceolate, margins entire, apex acute; petiole 2–11 cm, usually sparsely pubescent; blade unlobed, reniform to ovate, 2–4 × 2–4 cm, base cordate, lobes often overlapping, margins serrate, ciliate or eciliate, apex rounded or acute to acuminate, surfaces sometimes glabrous, usually sparsely pubescent adaxially. |
Peduncles | 2–12.5 cm, glabrous or puberulent. |
3–11 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm; petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins mostly eciliate, auricles 1–2 mm; petals white on both surfaces, lower 3 purple-veined, lateral 2 usually beardless, lowest 8–10 mm, spur white, gibbous, 1–2 mm; style head beardless; cleistogamous flowers axillary. |
Capsules | spherical to oblong, 5–12 mm, glabrous. |
ovoid to ellipsoid, 4–6 mm, glabrous. |
Seeds | light brown, 2.8–3.3 mm. |
beige to bronze, 1.5–2 mm. |
2n | = 24, 48. |
= 44, 48. |
Viola douglasii |
Viola blanda |
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Phenology | Flowering Feb–Jul. | Flowering Apr–Jun. |
Habitat | Vernally moist grassy slopes and flats, often serpentine soil (except Oregon) | Rich woods |
Elevation | 20–2300 m (100–7500 ft) | 30–2000 m (100–6600 ft) |
Distribution |
CA; OR; Mexico (Baja California)
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AL; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; NC; ND; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; AB; MB; NB; NL; NS; ON; PE; QC; SK
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Discussion | Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola blanda occurs in small colonies; individual plants are interconnected by stolons. Whether to recognize Viola incognita at any taxonomic level is currently unresolved. It is said to have pubescent leaf blades, greenish peduncles, nontwisted lateral petals, and a preference for moister habitats. Most of these characters fall within the range of variation observed in V. blanda. N. H. Russell (1965) noted that Viola incognita is principally found in glaciated areas whereas V. blanda is found in nonglaciated areas. Á. Löve and D. Löve (1982b) and J. M. Canne (1987) reported a chromosome count of 2n = 44; J. Clausen (1929) and A. Gershoy (1934) reported 2n = 48. V. B. Baird (1942) reported that V. blanda (and V. incognita) have fragrant flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 124. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. chrysantha | V. blanda var. palustriformis, V. incognita, V. incognita var. forbesii, V. leconteana |
Name authority | Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841) | Willdenow: Hort. Berol. 1(2): plate 24. (1804) |
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