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northern two-eyed or wedge-leaf violet, northern two-eyed violet, wedge-leaf violet

alpine marsh violet, marsh violet, northern marsh or marsh or swamp or alpine marsh violet, northern marsh violet, violette des marais

Habit Plants perennial, caulescent, not stoloniferous, 2–25 cm. Plants perennial, acaulescent, stoloniferous, 3–21.5 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy.
Stems

1–3, usually erect, sometimes prostrate or ascending, glabrous, from shallow, fleshy rhizome or deep-seated caudex with fleshy roots.

Leaves

basal and cauline;

basal: 2–6;

stipules lanceolate, margins entire or gland-toothed, apex acute;

petiole 4.5–9.8 cm, glabrous;

blade purple-veined, orbiculate-ovate to deltate, 1–4 × 2.7–3.8 cm, usually shiny, leathery, base cuneate, margins serrate, eciliate, apex acute, mucronulate, surfaces glabrous;

cauline similar to basal except: stipules lanceolate to ovate, margins entire or erose to lacerate, with or without gland-tipped processes, apex acute to acuminate;

petiole 0.5–20 cm;

blade usually rhombic, sometimes orbiculate, 0.9–2.6 × 0.7–1.8 cm, margins crenate to ± serrate, apex acute to obtuse.

basal, 2–4, ascending to erect;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1–17 cm, glabrous;

blade unlobed, reniform or ovate to orbiculate, 0.5–6.4 × 0.5–5.5 cm, ± as long as wide, base cordate, margins crenulate, eciliate, apex obtuse to acuminate, surfaces glabrous.

Peduncles

1–10.5 cm, glabrous.

2–20.7 cm, glabrous, bracteoles usually below middle in chasmogamous flowers.

Flowers

sepals lanceolate, margins eciliate, auricles 0.5–1 mm;

petals deep reddish violet abaxially, white adaxially, upper 2 sometimes with purple patch basally, lateral 2 with purple patch basally beyond smaller yellow area, usually bearded, sometimes beardless, lowest purple-veined with yellow area basally, 8–14 mm, spur yellow, gibbous, 1–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals ovate or lanceolate, margins eciliate, auricles 1–2 mm;

petals lilac, pale blue, or pale violet on both surfaces, sometimes nearly white and tinged with lilac, lowest and sometimes lateral 2 dark violet-veined, lateral 2 bearded or beardless, lowest 8–16 mm, spur usually same color as petals, gibbous, 1–3 mm;

style head beardless; cleistogamous flowers on ascending peduncles.

Capsules

± spherical, 5–9 mm, glabrous.

ellipsoid, 6–10 mm, glabrous.

Seeds

deep brown-purple, 2.1–3 mm.

dark brown, 1–2 mm.

2n

= 48.

Viola cuneata

Viola palustris

Phenology Flowering Mar–Sep. Flowering Apr–Jul.
Habitat Open pine or oak forests, often serpentine soil Marshes, swamps, fens, wet forests, stream banks, brushy places, lowlands to subalpine zones
Elevation 600–2200 m (2000–7200 ft) 0–1800 m (0–5900 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; ME; MT; NH; NV; OR; SD; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola cuneata often occurs on serpentine-derived soil and is closely related to V. flettii, V. hallii, and V. ocellata (V. B. Baird 1942). Originally described as beardless by Watson, the lateral petals of flowers in the same population can be prominently bearded or essentially beardless. The cauline blades of Viola cuneata are often vertical, especially early in season.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola palustris occurs in small colonies; individual plants are interconnected by stolons. When flowering, usually three or more leaves are present per caudex.

M. Sorsa (1968) noted that in western North America Viola palustris specimens differ from European V. palustris plants and often show variation in character combinations. She stated (pers. comm.) that North American V. palustris is not a uniform entity.

E. Hultén (1968) synonymized Viola palustris with V. epipsila, as did W. J. Cody (2000). G. W. Douglas et al. (1998–2002, vol. 5) recognized both species. We assume that V. palustris does not occur in Alaska; verification is needed. See additional discussion under 18a. V. epipsila var. repens.

M. Sorsa (1968) indicated that Viola palustris var. brevipes has often been confused with V. blanda and that var. brevipes may have arisen from hybridization between a tetraploid and a diploid plant. Listed here in synonymy, var. brevipes may warrant recognition.

Reports of Viola palustris from one location in Coconino County, Arizona, first collected in 1962, likely refer to V. macloskeyi.

Viola palustris hybridizes with V. epipsila var. repens (M. Sorsa 1968; T. Marcussen and T. Karlsson 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 129. FNA vol. 6, p. 142.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. palustris subsp. brevipes, V. palustris var. brevipes, V. palustris var. leimonia, V. palustris var. pensylvanica, V. palustris subsp. pubifolia
Name authority S. Watson: Proc. Amer. Acad. Arts 14: 290. (1879) Linnaeus: Sp. Pl. 2: 934. (1753)
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