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northern two-eyed or wedge-leaf violet, northern two-eyed violet, wedge-leaf violet

rock violet

Habit Plants perennial, caulescent, not stoloniferous, 2–25 cm. Plants perennial, caulescent, not stoloniferous, 5–15 cm.
Stems

1–3, usually erect, sometimes prostrate or ascending, glabrous, from shallow, fleshy rhizome or deep-seated caudex with fleshy roots.

1–3, ascending to erect, glabrous, on caudex from fleshy rhizome.

Leaves

basal and cauline;

basal: 2–6;

stipules lanceolate, margins entire or gland-toothed, apex acute;

petiole 4.5–9.8 cm, glabrous;

blade purple-veined, orbiculate-ovate to deltate, 1–4 × 2.7–3.8 cm, usually shiny, leathery, base cuneate, margins serrate, eciliate, apex acute, mucronulate, surfaces glabrous;

cauline similar to basal except: stipules lanceolate to ovate, margins entire or erose to lacerate, with or without gland-tipped processes, apex acute to acuminate;

petiole 0.5–20 cm;

blade usually rhombic, sometimes orbiculate, 0.9–2.6 × 0.7–1.8 cm, margins crenate to ± serrate, apex acute to obtuse.

basal and cauline;

basal: 1–4;

stipules unknown;

petiole 1–10 cm, glabrous, sometimes finely puberulent;

blade broadly ovate or deltate to broadly deltate, 1–2.5(–2.9) × 0.6–2.2(–2.6) cm, base usually cordate to truncate, rarely rounded, margins crenate-dentate, eciliate, apex acute, surfaces glabrous;

cauline similar to basal except: stipules lanceolate, margins usually fimbriate-toothed, sometimes entire, apex attenuate or acute;

petiole 1.1–3.7 cm, sometimes finely puberulent;

blade ovate to deltate, 0.7–2.2 × 0.4–1.2 cm, base sometimes rounded on distal blades.

Peduncles

1–10.5 cm, glabrous.

3–6(–10) cm, glabrous.

Flowers

sepals lanceolate, margins eciliate, auricles 0.5–1 mm;

petals deep reddish violet abaxially, white adaxially, upper 2 sometimes with purple patch basally, lateral 2 with purple patch basally beyond smaller yellow area, usually bearded, sometimes beardless, lowest purple-veined with yellow area basally, 8–14 mm, spur yellow, gibbous, 1–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate, margins eciliate, auricles 0.5–1 mm;

petals blue to pale violet on both surfaces with yellow area basally, lower 3 purple-veined, lowest with yellow area, lateral 2 bearded, lowest 5.5–11 mm, spur white to pale violet, gibbous, 0.5–1.3 mm;

style head bearded; cleistogamous flowers unknown.

Capsules

± spherical, 5–9 mm, glabrous.

subglobose, ca. 5 mm, glabrous.

Seeds

deep brown-purple, 2.1–3 mm.

dark brown, ca. 1.8 mm.

Viola cuneata

Viola lithion

Phenology Flowering Mar–Sep. Flowering Jun–Aug.
Habitat Open pine or oak forests, often serpentine soil Seasonally wet cracks and crevices, narrow ledges of rock outcrops, shaded northeast-facing avalanche chutes, cirque headwalls, subalpine conifer zone
Elevation 600–2200 m (2000–7200 ft) 2300–3100 m (7500–10200 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

Viola cuneata often occurs on serpentine-derived soil and is closely related to V. flettii, V. hallii, and V. ocellata (V. B. Baird 1942). Originally described as beardless by Watson, the lateral petals of flowers in the same population can be prominently bearded or essentially beardless. The cauline blades of Viola cuneata are often vertical, especially early in season.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Viola lithion is known only from the White Pine Range in Nevada and the Pilot Range straddling the Nevada-Utah border. It is related to V. canadensis and V. flettii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 129. FNA vol. 6, p. 136.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Name authority S. Watson: Proc. Amer. Acad. Arts 14: 290. (1879) N. H. Holmgren & P. K. Holmgren: Brittonia 44: 300, fig. 1A – D. (1992)
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