Viola cuneata |
Viola arvensis |
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northern two-eyed or wedge-leaf violet, northern two-eyed violet, wedge-leaf violet |
European field violet, European field-pansy, field pansy, field violet, pansy, violette des champs, wild pansy |
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Habit | Plants perennial, caulescent, not stoloniferous, 2–25 cm. | Plants annual, caulescent, not stoloniferous, 5–35 cm. |
Stems | 1–3, usually erect, sometimes prostrate or ascending, glabrous, from shallow, fleshy rhizome or deep-seated caudex with fleshy roots. |
1–5, usually erect, sometimes prostrate or decumbent, branched, subglabrous or puberulent, clustered on taproot. |
Leaves | basal and cauline; basal: 2–6; stipules lanceolate, margins entire or gland-toothed, apex acute; petiole 4.5–9.8 cm, glabrous; blade purple-veined, orbiculate-ovate to deltate, 1–4 × 2.7–3.8 cm, usually shiny, leathery, base cuneate, margins serrate, eciliate, apex acute, mucronulate, surfaces glabrous; cauline similar to basal except: stipules lanceolate to ovate, margins entire or erose to lacerate, with or without gland-tipped processes, apex acute to acuminate; petiole 0.5–20 cm; blade usually rhombic, sometimes orbiculate, 0.9–2.6 × 0.7–1.8 cm, margins crenate to ± serrate, apex acute to obtuse. |
cauline; stipules palmately lobed, middle lobe oblanceolate, obovate, elliptic, or lanceolate, ± equaling leaf blade, proximal lobes dissected, shorter, margins ciliate, apex acute to obtuse; petiole 0.5–2.3 cm, glabrous or puberulent; blade: proximal ovate to ± oblong, distal narrowly or broadly lanceolate, 0.8–3.4 × 0.3–1.9 cm, base attenuate to ± truncate, margins coarsely crenate-serrate, ciliate or eciliate, apex acute to obtuse, surfaces pubescent abaxially, at least on major veins, glabrous adaxially. |
Peduncles | 1–10.5 cm, glabrous. |
2–8 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep reddish violet abaxially, white adaxially, upper 2 sometimes with purple patch basally, lateral 2 with purple patch basally beyond smaller yellow area, usually bearded, sometimes beardless, lowest purple-veined with yellow area basally, 8–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles 2–4 mm; petals white to pale yellow on both surfaces, upper 2 ± violet, lower 3 with yellow basal area, often violet-veined, lateral 2 bearded, ± equaling or shorter than sepals, lowest with dark yellow area basally, 7–15 mm, spur blue-violet to purple, elongated, 3–5 mm; style head bearded; cleistogamous flowers absent. |
Capsules | ± spherical, 5–9 mm, glabrous. |
± spherical, 5–9 mm, glabrous. |
Seeds | deep brown-purple, 2.1–3 mm. |
brown, 1.5–1.9 mm. |
2n | = 34. |
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Viola cuneata |
Viola arvensis |
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Phenology | Flowering Mar–Sep. | Flowering May–Jul. |
Habitat | Open pine or oak forests, often serpentine soil | Abandoned fields, roadsides, lawns |
Elevation | 600–2200 m (2000–7200 ft) | 0–3000 m (0–9800 ft) |
Distribution |
CA; OR
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CA; CT; DC; DE; GA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; SPM; Asia (Siberia); Greenland; Europe; Africa [Introduced in North America]
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Discussion | Viola cuneata often occurs on serpentine-derived soil and is closely related to V. flettii, V. hallii, and V. ocellata (V. B. Baird 1942). Originally described as beardless by Watson, the lateral petals of flowers in the same population can be prominently bearded or essentially beardless. The cauline blades of Viola cuneata are often vertical, especially early in season. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A. R. Clapham et al. (1987) noted that Viola arvensis is pollinated by insects and is often selfed; T. Marcussen and T. Karlsson (2010) stated that V. arvensis regularly self-pollinates. Roots of V. arvensis have the odor of wintergreen when crushed (A. E. Radford et al. 1968; W. J. Hayden and J. Clough 1990). Viola arvensis is not vegetatively distinguishable from V. tricolor var. tricolor. G. Halliday (pers. comm.) reported that Viola arvensis occurs in southwestern Greenland, where it is introduced, and is ephemeral on St. Pierre and Miquelon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 129. | FNA vol. 6, p. 121. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | Mnemion arvense, V. tricolor var. arvensis | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 14: 290. (1879) | Murray: Prodr. Stirp. Gott., 73. (1770) |
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