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blue marsh violet, marsh blue or northern bog or marsh or blue marsh violet, marsh blue violet, violette cucullée

Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

Habit Plants perennial, acaulescent, not stoloniferous, 5–20 cm; rhizome thick, fleshy. Plants perennial, caulescent, not stoloniferous, 3–20 cm.
Stems

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

Leaves

basal, 4–8, ascending to erect;

stipules linear-lanceolate, margins entire, apex acute;

petiole 6–20 cm, usually glabrous;

blade unlobed, ovate to reniform, 1.5–11 × 1–9 cm, base reniform to cordate, margins crenate to serrate, ciliate, apex acute or obtuse to slightly caudate, surfaces glabrous, rarely pubescent.

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

Peduncles

7–25 cm, glabrous or pubescent.

2–12.5 cm, glabrous or puberulent.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 3–6 mm;

petals light to dark blue-violet to violet on both surfaces, upper 2 and lateral 2 darker basally, lateral 2 densely bearded, lowest white basally, dark purple-veined, 9–13 mm, spur color same as petals, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on erect peduncles.

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

Capsules

ellipsoid to ± ovoid, 10–15 mm, glabrous.

spherical to oblong, 5–12 mm, glabrous.

Seeds

beige, mottled to bronze, 1–2 mm.

light brown, 2.8–3.3 mm.

2n

= 54.

= 24, 48.

Viola cucullata

Viola douglasii

Phenology Flowering Apr–Jun. Flowering Feb–Jul.
Habitat Open or wooded wet areas, seeps, springs, swamps, marshes, streamsides Vernally moist grassy slopes and flats, often serpentine soil (except Oregon)
Elevation 0–2000 m (0–6600 ft) 20–2300 m (100–7500 ft)
Distribution
from FNA
AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; MS; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; NB; NF; NS; ON; PE; QC; SPM
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola cucullata reportedly hybridizes with V. sororia (= V. ×bissellii House), V. brittoniana (= V. ×notabilis E. P. Bicknell), and V. sagittata var. sagittata (= V. ×porteriana House).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 128. FNA vol. 6, p. 129.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. cucullata var. leptosepala, V. cucullata var. macrotis, V. cucullata var. microtitis, V. dicksonii, V. leptosepala, V. macrotis V. chrysantha
Name authority Aiton: Hort. Kew. 3: 288. (1789) Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841)
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