Viola bicolor
Viola palustris
field pansy, violette de Rafinesque, wild pansy
alpine marsh violet, marsh violet, northern marsh or marsh or swamp or alpine marsh violet, northern marsh violet, violette des marais
basal, 2–4, ascending to erect;
stipules linear-lanceolate, margins entire, apex acute;
petiole 1–17 cm, glabrous;
blade unlobed, reniform or ovate to orbiculate, 0.5–6.4 × 0.5–5.5 cm, ± as long as wide, base cordate, margins crenulate, eciliate, apex obtuse to acuminate, surfaces glabrous.
1–4.5 cm, glabrous or pubescent.
2–20.7 cm, glabrous, bracteoles usually below middle in chasmogamous flowers.
sepals ovate to lanceolate, margins ciliate or eciliate, auricles 0.5–2 mm;
petals white or cream to pale bluish violet on both surfaces, dark purple-veined, lateral 2 longer than sepals, bearded, lowest 8–10 mm, spur white to blue-violet, gibbous, 1–1.5 mm, shorter than or equaling sepal auricles;
style head bearded; cleistogamous flowers axillary.
sepals ovate or lanceolate, margins eciliate, auricles 1–2 mm;
petals lilac, pale blue, or pale violet on both surfaces, sometimes nearly white and tinged with lilac, lowest and sometimes lateral 2 dark violet-veined, lateral 2 bearded or beardless, lowest 8–16 mm, spur usually same color as petals, gibbous, 1–3 mm;
style head beardless; cleistogamous flowers on ascending peduncles.
ellipsoid to oblong, 4–7 mm, glabrous.
ellipsoid, 6–10 mm, glabrous.
beige to bronze, 0.3–1.5 mm.
dark brown, 1–2 mm.
= 34.
= 48.
Viola bicolor
Viola palustris
Viola bicolor is the only pansy native to North America (V. B. Baird 1942; J. Clausen et al. 1964; A. E. Radford et al. 1968) and is the only annual Viola species that produces cleistogamous flowers (Baird; A. Gershoy 1934). Roots of V. bicolor have the odor of wintergreen when crushed (W. J. Hayden and J. Clough 1990).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Viola palustris occurs in small colonies; individual plants are interconnected by stolons. When flowering, usually three or more leaves are present per caudex.
M. Sorsa (1968) noted that in western North America Viola palustris specimens differ from European V. palustris plants and often show variation in character combinations. She stated (pers. comm.) that North American V. palustris is not a uniform entity.
E. Hultén (1968) synonymized Viola palustris with V. epipsila, as did W. J. Cody (2000). G. W. Douglas et al. (1998–2002, vol. 5) recognized both species. We assume that V. palustris does not occur in Alaska; verification is needed. See additional discussion under 18a. V. epipsila var. repens.
M. Sorsa (1968) indicated that Viola palustris var. brevipes has often been confused with V. blanda and that var. brevipes may have arisen from hybridization between a tetraploid and a diploid plant. Listed here in synonymy, var. brevipes may warrant recognition.
Reports of Viola palustris from one location in Coconino County, Arizona, first collected in 1962, likely refer to V. macloskeyi.
Viola palustris hybridizes with V. epipsila var. repens (M. Sorsa 1968; T. Marcussen and T. Karlsson 2010).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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