Viola beckwithii
Viola sheltonii
Beckwith's or Great Basin violet, Beckwith's violet, Great Basin violet, sagebrush pansy, sagebrush violet
cut-leaf violet, fan violet, Shelton's violet
1–3, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous or usually puberulent, on single, short, vertical, deep-seated caudex.
1–3, prostrate, decumbent, or erect, glabrous or sparsely puberulent, from short, often vertical, deep-seated or usually shallow, subligneous rhizome.
basal and cauline;
basal: 1–6 per caudex, palmately compound, ± 2-ternate or 3-ternate, leaflets 3;
stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute;
petiole 2–10.5 cm, usually puberulent;
blade ovate to deltate, 2.4–5 × 3.5–4.5 cm, base tapered, ultimate leaflets dissected into oblong, elliptic, lanceolate, or oblanceolate lobes 1–7 mm wide, lobe margins entire, usually ciliate, apex acute to obtuse, mucronulate, surfaces usually puberulent, seldom glabrous, abaxial surface without prominent vein parallel to each margin;
cauline similar to basal except: stipules linear, apex acuminate;
petiole 2–5.7 cm, usually puberulent, rarely glabrous;
blade 1–2.7 × 1.5–3 cm.
basal and cauline;
basal: 1–3, palmately compound, leaflets 3;
stipules lanceolate-ovate, margins laciniate with gland-tipped projections, apex acute to acuminate;
petiole 8.6–21 cm, glabrous or sparsely puberulent;
blade reniform or ovate to ± orbiculate, 2–7 × 2–11 cm, coriaceous, base tapered, each leaflet cleft or dissected into 3 ± obovate lobes, each lobe further divided into 2–3 oblanceolate, pandurate, spatulate, oblong, lanceolate, or elliptic, lobes 2–10 mm wide, margins entire, ciliate or eciliate, apex acute to obtuse, mucronulate, surfaces glabrous or sparsely puberulent;
cauline similar to basal except: stipules ovate to lanceolate, margin projections gland-tipped or eglandular, apex long-acuminate;
petiole 5.5–12 cm;
blade 1.2–6.3 × 1.2–10.5 cm.
1.5–10.6(–15.7) cm, usually puberulent, seldom glabrous.
5–19 cm, glabrous or sparsely puberulent.
sepals lanceolate, margins eciliate, auricles 0.1–1 mm;
petals dark reddish violet on both surfaces, lower 3 usually lilac, rarely white or whitish, lateral 2 bearded, with yellow patch basally, dark reddish violet-veined, lowest 10–22 mm, with yellow patch, dark reddish violet-veined, spur whitish or yellowish, tinged purple, gibbous, 0.5–2 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 dark brown to brownish purple abaxially, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded or beardless, lowest 7–18 mm, spur yellowish with brownish purple specks, gibbous, 1–2 mm;
style head bearded; cleistogamous flowers axillary.
oblong-ovoid, 7–12 mm, glabrous.
oblong to ovoid, 6–8 mm, glabrous or puberulent.
brown, 3–4 mm.
brownish, shiny, ca. 2.5 mm.
= 24.
= 12.
Viola beckwithii
Viola sheltonii
In some populations of Viola beckwithii, the three lower petals are white with a yellow area proximally (V. B. Baird 1942). Leaves have been described as palmately biternate or triternate (L. Abrams and R. S. Ferris 1923–1960, vol. 3), ternately decompound into linear segments (C. L. Hitchcock et al. 1955–1969, vol. 3), palmately three-parted then bipinnately parted into ultimate linear or spatulate segments (P. A. Munz 1959), and palmately about three times three-parted into linear or spatulate-linear segments (W. L. Jepson 1951). Some populations in northern California are nearly or completely glabrous, which M. S. Baker recognized as var. glabrata.
Viola beckwithii is reported to hybridize with V. utahensis (G. Davidse 1976). Observed pollinators of V. beckwithii in Utah include Apis mellifera Linnaeus and Anthophora ursina Cresson (Davidse).
When Cottam described Viola bonnevillensis, he suggested that it could be a hybrid between V. beckwithii and V. utahensis, and G. Davidse (1976) concurred. The type specimen of V. bonnevillensis (Cottam 7067, UT) was examined by R. J. Little. Because no similar forms are known to have been collected since 1939, it is presumed that this taxon is a hybrid.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The cleistogamous flowers of Viola sheltonii are borne on long, prostrate peduncles usually buried in duff around the plant. Mature cleistogamous capsules are usually hidden and the dehisced seeds remain close to the parent plant. Some populations of V. sheltonii produce only cleistogamous flowers (D. Klaber 1976).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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