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Beckwith's or Great Basin violet, Beckwith's violet, Great Basin violet, sagebrush pansy, sagebrush violet

primrose-leaf violet, western bog violet, western white bog violet

Habit Plants perennial, caulescent, not stoloniferous, 2–22 cm. Plants perennial, acaulescent, stoloniferous, (3–)5–20(–36) cm; stolons pale, often rooting and leafy at nodes; rhizome thick or slender, fleshy.
Stems

1–3, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous or usually puberulent, on single, short, vertical, deep-seated caudex.

Leaves

basal and cauline;

basal: 1–6 per caudex, palmately compound, ± 2-ternate or 3-ternate, leaflets 3;

stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute;

petiole 2–10.5 cm, usually puberulent;

blade ovate to deltate, 2.4–5 × 3.5–4.5 cm, base tapered, ultimate leaflets dissected into oblong, elliptic, lanceolate, or oblanceolate lobes 1–7 mm wide, lobe margins entire, usually ciliate, apex acute to obtuse, mucronulate, surfaces usually puberulent, seldom glabrous, abaxial surface without prominent vein parallel to each margin;

cauline similar to basal except: stipules linear, apex acuminate;

petiole 2–5.7 cm, usually puberulent, rarely glabrous;

blade 1–2.7 × 1.5–3 cm.

basal, 4–8, prostrate to ascending;

stipules linear-lanceolate, margins ± crenate-serrate, sometimes glandular, apex acute;

petiole (1–)3–13(–29) cm, glabrous or pubescent;

blade unlobed, elliptic to narrowly or broadly ovate, (1.5–)3–7(–9) × 1–3(–3.5) cm, longer than wide, base broadly cordate to attenuate, rarely ± truncate, margins crenulate to serrulate, sometimes glandular, mostly eciliate, apex acute to rounded, mucronulate, surfaces glabrous or sparsely pubescent.

Peduncles

1.5–10.6(–15.7) cm, usually puberulent, seldom glabrous.

(3–)6–18(–28) cm, glabrous.

Flowers

sepals lanceolate, margins eciliate, auricles 0.1–1 mm;

petals dark reddish violet on both surfaces, lower 3 usually lilac, rarely white or whitish, lateral 2 bearded, with yellow patch basally, dark reddish violet-veined, lowest 10–22 mm, with yellow patch, dark reddish violet-veined, spur whitish or yellowish, tinged purple, gibbous, 0.5–2 mm;

style head bearded; cleistogamous flowers absent.

sepals lanceolate to ovate, margins usually eciliate, auricles 1–2 mm;

petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded or beardless, lowest 9–14(–16) mm, spur white, gibbous, 1–2 mm;

style head beardless; cleistogamous flowers present or absent.

Capsules

oblong-ovoid, 7–12 mm, glabrous.

ellipsoid, (5–)8–9 mm, glabrous.

Seeds

brown, 3–4 mm.

beige to bronze, 1.5–2 mm.

2n

= 24.

= 24.

Viola beckwithii

Viola primulifolia

Phenology Flowering Mar–May.
Habitat Dry or moist places, among shrubs or beneath pines
Elevation 900–2700 m (3000–8900 ft)
Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IL; IN; KY; LA; MA; MD; ME; MI; MN; MS; NC; NH; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; VT; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

In some populations of Viola beckwithii, the three lower petals are white with a yellow area proximally (V. B. Baird 1942). Leaves have been described as palmately biternate or triternate (L. Abrams and R. S. Ferris 1923–1960, vol. 3), ternately decompound into linear segments (C. L. Hitchcock et al. 1955–1969, vol. 3), palmately three-parted then bipinnately parted into ultimate linear or spatulate segments (P. A. Munz 1959), and palmately about three times three-parted into linear or spatulate-linear segments (W. L. Jepson 1951). Some populations in northern California are nearly or completely glabrous, which M. S. Baker recognized as var. glabrata.

Viola beckwithii is reported to hybridize with V. utahensis (G. Davidse 1976). Observed pollinators of V. beckwithii in Utah include Apis mellifera Linnaeus and Anthophora ursina Cresson (Davidse).

When Cottam described Viola bonnevillensis, he suggested that it could be a hybrid between V. beckwithii and V. utahensis, and G. Davidse (1976) concurred. The type specimen of V. bonnevillensis (Cottam 7067, UT) was examined by R. J. Little. Because no similar forms are known to have been collected since 1939, it is presumed that this taxon is a hybrid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Viola primulifolia occurs in small colonies; individual plants are interconnected by stolons.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades elliptic to widely ovate, base broadly cordate to attenuate, surfaces sparsely pubescent, rarely glabrous; cleistogamous flowers present; c, e United States.
var. primulifolia
1. Leaf blades elliptic to narrowly ovate, base usually attenuate, rarely ± truncate, surfaces glabrous, rarely pubescent; cleistogamous flowers absent; nw California, sw Oregon.
var. occidentalis
Source FNA vol. 6, p. 122. FNA vol. 6, p. 148.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. primulifolia var. occidentalis, V. primulifolia var. primulifolia
Synonyms V. beckwithii var. cachensis, V. beckwithii subsp. glabrata, V. bonnevillensis
Name authority Torrey: in E. M. Durand, Pl. Pratten. Calif., 82. (1855) Linnaeus: Sp. Pl. 2: 934. (1753)
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