Viola bakeri
Viola palmata
Baker's violet, yellow prairie violet
early blue violet, three-lobe violet, three-lobed violet, trilobed violet, wood violet
1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome.
basal and cauline;
basal: 1–4;
stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate;
petiole 1–15.4 cm, glabrous or puberulent;
blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins;
cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections;
petiole 1.5–7.5 cm;
blade 1.9–6.7 × 0.5–1.6 cm.
basal, 2–3, ascending to erect;
stipules linear-lanceolate, margins entire, apex acute;
petiole 1–20 cm, glabrous or pubescent;
earliest leaf blades unlobed, mid-season blades 3–9-lobed, lobes sometimes appearing petiolate and sometimes further lobed;
earliest leaf blades reniform to ovate, mid-season blades with middle lobes usually ovate or elliptic to widely obovate, sometimes narrowly elliptic, narrowly ovate, lanceolate, or spatulate to narrowly obovate, lateral lobes elliptic, obdeltate, or spatulate to falcate, smaller lobes similar, 1–14 × 1–10 cm, base truncate to cordate, margins serrate or entire, usually ciliate, apex acute, rounded, blunt, or obtuse, surfaces glabrous or pubescent throughout or along veins.
1.5–11.6 cm, glabrous or pubescent.
3–13 cm, glabrous or pubescent.
sepals lanceolate, margins eciliate, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm;
style head bearded; cleistogamous flowers axillary.
sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm;
petals violet on both surfaces, lower 3 white basally, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, spur sometimes bearded, lowest 15–25 mm, spur white, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent.
ellipsoid, 5–15 mm, glabrous.
light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus.
beige, mottled to bronze, 1.5–2.5 mm.
= 48.
= 54.
Viola bakeri
Viola palmata
M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3).
Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri
from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
N. L. Gil-Ad (1997) presented evidence based on seed coat micromorphology that some plants known as Viola palmata are hybrids between plants with lobed and unlobed leaves and believed the type specimen to be hybrid in origin. He chose not to recognize the name V. palmata, choosing instead to recognize V. triloba. If the type specimen does represent a hybrid, Gil-Ad would be correct in recognizing V. triloba. Because the purity of the type specimen cannot be ascertained, we consider V. palmata the most appropriate name for this taxon. We acknowledge the presence of hybrids between lobed and unlobed plants; such hybrids obscure lines of demarcation. Some herbarium specimens may represent such hybrids; we feel most do not.
L. E. McKinney (1992) described the nomenclatural history of Viola palmata. Homophylly versus heterophylly differentiates V. palmata from V. subsinuata, the homophyllous violet most often called V. palmata. We know that the taxon described by Linnaeus was heterophyllous, with some undivided leaf blades.
Varieties 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Petioles, leaf surfaces, and peduncles usually pubescent, rarely glabrous; apex of middle leaf lobe acute; plants of dry to mesic habitats. | var. palmata |
1. Petioles, leaf surfaces, and peduncles usually glabrous, rarely pubescent; apex of middle leaf lobe rounded, blunt, or obtuse; plants of wet ground. | var. heterophylla |
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