Viola bakeri |
Viola missouriensis |
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Baker's violet, yellow prairie violet |
Missouri violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–30 cm. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizomes thick, fleshy. |
Stems | 1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–4; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate; petiole 1–15.4 cm, glabrous or puberulent; blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins; cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections; petiole 1.5–7.5 cm; blade 1.9–6.7 × 0.5–1.6 cm. |
basal, 1–8, ascending to erect; stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes distally glandular, apex acute; petiole 5–20 cm, glabrous; blade green abaxially, unlobed, usually narrowly to broadly deltate, 1.5–12 × 1.5–10 cm, not fleshy, base cordate or broadly cordate to truncate, margins ± crenate to serrate, ciliate or eciliate, apex acute to acuminate, surfaces glabrous, rarely pubescent. |
Peduncles | 1.5–11.6 cm, glabrous or pubescent. |
3–25 cm, usually glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest beardless, rarely lightly bearded, 15–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent. |
ellipsoid, 5–12 mm, glabrous. |
Seeds | light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 48. |
= 54. |
Viola bakeri |
Viola missouriensis |
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Phenology | Flowering May–Jul. | Flowering Mar–May. |
Habitat | Wet and dry places in openings of coniferous forests | Swamps, thickets, stream banks, alluvial woods |
Elevation | 900–3800 m (3000–12500 ft) | 50–2000 m (200–6600 ft) |
Distribution |
CA; NV; OR; WA
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AL; AR; DC; DE; FL; GA; IA; IL; IN; KS; LA; MD; MN; MO; MS; NC; NE; NJ; NM; OK; SC; SD; TN; TX; VA; WI
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Discussion | M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3). Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
What many have treated as Viola affinis, especially in the southern Gulf coastal states, is likely to be V. missouriensis. N. H. Russell (1965) considered the two as likely part of a species complex inhabiting alluvial woods and wet areas and exhibiting the typical deltate leaf blade shape. L. E. McKinney (1992) considered V. missouriensis a variety of V. sororia. Viola missouriensis appears to have a closer affinity to V. affinis, as Russell suggested; current evidence suggests maintaining V. missouriensis as a separate species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 121. | FNA vol. 6, p. 138. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. bakeri subsp. shastensis, V. nuttallii var. bakeri | V. candidula, V. langloisii, V. lucidifolia, V. sororia var. missouriensis |
Name authority | Greene: Pittonia 3: 307. (1898) | Greene: Pittonia 4: 141. (1900) |
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