Viola bakeri
Viola macloskeyi
Baker's violet, yellow prairie violet
Macloskey's or northern white or small white or smooth white or wild white violet, Macloskey's violet, northern white violet, small white violet, smooth white violet, violette pâle
1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome.
basal and cauline;
basal: 1–4;
stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate;
petiole 1–15.4 cm, glabrous or puberulent;
blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins;
cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections;
petiole 1.5–7.5 cm;
blade 1.9–6.7 × 0.5–1.6 cm.
basal, 2–6, ascending to erect;
stipules ovate to linear-lanceolate, margins entire or glandular-toothed, apex acute;
petiole 1–10 cm, strigose;
blade unlobed, reniform to ovate, 1–6.5 × 1–5.5 cm, base broadly or shallowly cordate, margins ± entire or shallowly crenate, eciliate, apex rounded to acute, surfaces usually glabrous, abaxial surfaces sometimes puberulent on proximal 1/2.
1.5–11.6 cm, glabrous or pubescent.
2.5–11(–21) cm, usually glabrous, sometimes villous.
sepals lanceolate, margins eciliate, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm;
style head bearded; cleistogamous flowers axillary.
sepals lanceolate to ovate, margins eciliate, auricles 0.5–2 mm;
petals white on both surfaces, lower 3 purple-veined, lateral 2 bearded, rarely beardless, lowest 6–12 mm, spur white, gibbous, 1–2.5 mm;
style head beardless; cleistogamous flowers on ascending peduncles.
spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent.
ovoid, 5–9 mm, glabrous.
light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus.
beige to bronze, 1–1.5 mm.
= 48.
= 24.
Viola bakeri
Viola macloskeyi
M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3).
Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri
from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Viola macloskeyi was described by Banks in 1824 as V. rotundifolia var. pallens, from specimens collected by Banks in Labrador. E. Brainerd made the combination V. pallens in 1905 and later (1924) detailed its troublesome nomenclatural history.
Viola macloskeyi was described in 1895 by F. E. Lloyd based on specimens collected at the base of Mount Hood, Oregon. M. S. Baker (1953) placed V. macloskeyi as a subspecies of V. pallens. He later (1953b) corrected that change based on priority and thus the taxon became known as V. macloskeyi subsp. pallens. N. H. Russell (1955) maintained that status, separating subspp. macloskeyi and pallens on the wider, spreading basal leaf lobes of subsp. pallens. McKinney, after examining specimens of both taxa, including types, concluded that the differences have been exaggerated and fall within the range of variation of a single species.
Viola macloskeyi occurs in small colonies; individual plants are interconnected by stolons.
In parts of its range, Viola macloskeyi often shares habitat with V. renifolia; it may be difficult to distinguish the two. The stolons of V. macloskeyi are useful for identification during summer months; V. renifolia plants lack stolons. Patterns of indument can be useful; V. renifolia is usually hairy and V. macloskeyi is usually glabrous (petioles sometimes pubescent).
Viola macloskeyi reportedly hybridizes with V. primulifolia var. primulifolia (= V. ×mollicula House).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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