Viola bakeri |
Viola labradorica |
|
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Baker's violet, yellow prairie violet |
alpine violet, American dog or Labrador or alpine violet, American dog violet, dog violet, Labrador violet, violette du Labrador |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–30 cm. | Plants perennial, caulescent, not stoloniferous, 5–20 cm. |
Stems | 1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome. |
1–5, erect (later reclining), glabrous, on caudex from subligneous rhizome. |
Leaves | basal and cauline; basal: 1–4; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate; petiole 1–15.4 cm, glabrous or puberulent; blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins; cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections; petiole 1.5–7.5 cm; blade 1.9–6.7 × 0.5–1.6 cm. |
basal and cauline; basal: 1–5; stipules mostly lanceolate-linear, margins usually laciniate with gland-tipped projections, apex acute; petiole 1–9.5 cm, glabrous or puberulent; blade reniform or ovate to deltate, 1.4–5.5 × 1.8–5.3 cm, base cordate, not decurrent on petiole, margins crenate or serrate, mostly eciliate, apex acute to obtuse, surfaces usually glabrous, sometimes pubescent adaxially near margin; cauline similar to basal except: stipules lanceolate to ovate, margins ± entire or laciniate, projections often gland-tipped, apex acuminate; petiole 0.5–3.6 cm, glabrous; blade narrowly ovate to orbiculate, 1.8–2.9 × 1.5–3.1 cm, base deeply cordate to truncate, margins regularly crenate, apex obtuse to broadly rounded or apiculate. |
Peduncles | 1.5–11.6 cm, glabrous or pubescent. |
5–9 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 1–1.5 mm; petals usually lavender-violet to violet on both surfaces, rarely white, lower 3 white basally, darker violet-veined, lateral 2 sparsely bearded, lowest 9–16 mm, spur white to pinkish violet, elongated, 4–8 mm, tip usually straight, sometimes curved up; style head beardless; cleistogamous flowers unknown. |
Capsules | spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent. |
ovoid to ellipsoid, 5–7 mm, glabrous. |
Seeds | light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus. |
beige to bronze, 1.5–2 mm. |
2n | = 48. |
= 20. |
Viola bakeri |
Viola labradorica |
|
Phenology | Flowering May–Jul. | Flowering Apr–Aug. |
Habitat | Wet and dry places in openings of coniferous forests | Rich, mesic to wet woods, swamps, bogs |
Elevation | 900–3800 m (3000–12500 ft) | 50–3000 m (200–9800 ft) |
Distribution |
CA; NV; OR; WA
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AL; CO; CT; DC; DE; GA; IL; IN; KY; MA; MD; ME; MI; MN; NC; ND; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; MB; NB; NL; NS; NT; ON; PE; QC; SK; SPM; Greenland
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Discussion | M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3). Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. E. Ballard (1994) made a compelling case for including Viola conspersa within V. labradorica, which is followed here. Ballard (1992) did not recognize V. labradorica as occurring in Colorado. Several western herbaria have collections identified as V. labradorica (ASC, CS, KHD, RM, RMBL; http://swbiodiversity.org/seinet/index.pho). Labels on some collections at KHD imply that V. labradorica is considered synonymous with V. adunca var. bellidifolia. Viola labradorica reportedly hybridizes with V. adunca, V. rostrata (= V. ×malteana House), and V. striata (= V. ×eclipes H. E. Ballard), and less frequently with V. walteri var. appalachiensis and var. walteri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 121. | FNA vol. 6, p. 135. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. bakeri subsp. shastensis, V. nuttallii var. bakeri | V. adunca var. minor, V. canina var. muhlenbergii, V. conspersa, V. leucopetala, V. muhlenbergiana, V. muhlenbergiana var. minor, V. muhlenbergii |
Name authority | Greene: Pittonia 3: 307. (1898) | Schrank: Denkschr. Kõnigl.-Baier. Bot. Ges. Regensburg 1(2): 12. (1818) |
Web links |