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Baker's violet, yellow prairie violet

Howell's violet, violette de Howell

Habit Plants perennial, caulescent, not stoloniferous, 3–30 cm. Plants perennial, caulescent, not stoloniferous, 2–44 cm.
Stems

1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome.

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

Leaves

basal and cauline;

basal: 1–4;

stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate;

petiole 1–15.4 cm, glabrous or puberulent;

blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins;

cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections;

petiole 1.5–7.5 cm;

blade 1.9–6.7 × 0.5–1.6 cm.

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

Peduncles

1.5–11.6 cm, glabrous or pubescent.

2.8–17.8 cm, glabrous or sparsely pubescent.

Flowers

sepals lanceolate, margins eciliate, auricles 0.5–1 mm;

petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm;

style head bearded; cleistogamous flowers axillary.

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

Capsules

spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent.

ellipsoid, 7–11 mm, glabrous.

Seeds

light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus.

light brown, 2.1–2.8 mm.

2n

= 48.

= 40, 80.

Viola bakeri

Viola howellii

Phenology Flowering May–Jul. Flowering Apr–Jul.
Habitat Wet and dry places in openings of coniferous forests Moist, shady areas, coniferous forests, prairies, along streams
Elevation 900–3800 m (3000–12500 ft) 50–1500 m (200–4900 ft)
Distribution
from FNA
CA; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3).

Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri

from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 121. FNA vol. 6, p. 134.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. bakeri subsp. shastensis, V. nuttallii var. bakeri
Name authority Greene: Pittonia 3: 307. (1898) A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887)
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