Viola bakeri
Viola douglasii
Baker's violet, yellow prairie violet
Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet
1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome.
1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.
basal and cauline;
basal: 1–4;
stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate;
petiole 1–15.4 cm, glabrous or puberulent;
blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins;
cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections;
petiole 1.5–7.5 cm;
blade 1.9–6.7 × 0.5–1.6 cm.
basal and cauline;
basal: 1–6, bipinnately compound, leaflets 3–5;
stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;
petiole 5–6.8 cm, glabrous or puberulent;
blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;
cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;
petiole 0.9–4 cm;
blade 1.1–4.1 × 1–3.6 cm.
1.5–11.6 cm, glabrous or pubescent.
2–12.5 cm, glabrous or puberulent.
sepals lanceolate, margins eciliate, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm;
style head bearded; cleistogamous flowers axillary.
sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;
petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;
style head bearded; cleistogamous flowers absent.
spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent.
spherical to oblong, 5–12 mm, glabrous.
light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus.
light brown, 2.8–3.3 mm.
= 48.
= 24, 48.
Viola bakeri
Viola douglasii
M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3).
Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri
from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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