Viola bakeri |
Viola affinis |
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Baker's violet, yellow prairie violet |
Le Conte's violet, sand violet, violette affine |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–30 cm. | Plants perennial, acaulescent, not stoloniferous, 5–15 cm; rhizome slender, becoming thick and fleshy with age. |
Stems | 1–4, usually erect, sometimes prostrate or decumbent, leafy proximally and distally, usually puberulent, from usually vertical, subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–4; stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire, apex of each wing free, acute to acuminate; petiole 1–15.4 cm, glabrous or puberulent; blade lanceolate, oblanceolate, or elliptic, rarely ovate, 1.8–8.8 × 0.7–3.9 cm, thin, base attenuate, often oblique, margins usually entire, sometimes with a few sharp teeth or crenulate, ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent on margins or veins; cauline similar to basal except: stipules ovate to lanceolate, margins entire or lacerate, sometimes with glandular projections, apex with 2–3 projections; petiole 1.5–7.5 cm; blade 1.9–6.7 × 0.5–1.6 cm. |
basal, 1–6, ascending to erect; stipules lanceolate, margins entire or fimbriate, apex acute; petiole 2–10 cm, glabrous; blade green abaxially, unlobed, narrowly to broadly ovate or narrowly deltate, 1.5–10 × 1.5–10 cm, not fleshy, base cordate to broadly cordate or almost truncate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse, surfaces sparsely pubescent adaxially, rarely glabrous. |
Peduncles | 1.5–11.6 cm, glabrous or pubescent. |
3–15 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 sparsely bearded, lowest 6–14 mm, spur yellow, gibbous, 1–2 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals lavender-violet to dull reddish violet on both surfaces, lower 3 white basally and darker violet-veined, lateral 2 bearded, lowest 10–22 mm, usually obviously bearded, rarely beardless, spur white or same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers from prostrate to ascending peduncles. |
Capsules | spherical to ovoid, 5–10 mm, usually glabrous, rarely finely puberulent. |
often reddish or purplish-flecked or green, ellipsoid, 5–10 mm, glabrous or puberulent. |
Seeds | light to medium brown or dark red-brown, 2.6–3.1 mm, elaiosome not covering funiculus. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 48. |
= 54. |
Viola bakeri |
Viola affinis |
|
Phenology | Flowering May–Jul. | Flowering Apr–Jun. |
Habitat | Wet and dry places in openings of coniferous forests | Open or wooded wet areas, meadows, stream banks, thickets, shores of lakes, seasonally dry areas. |
Elevation | 900–3800 m (3000–12500 ft) | 100–2000 m (300–6600 ft) |
Distribution |
CA; NV; OR; WA
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CT; DC; DE; IL; IN; MA; MD; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; ON; QC |
Discussion | M. S. Baker (1957) wrote that Viola bakeri flowers were often without a brownish tinge on the back. Subsequent collections have shown that the upper two petals of V. bakeri are often brownish purple abaxially and thus this characteristic cannot be used to distinguish V. bakeri from V. praemorsa, as previously done (P. A. Munz 1959; C. L. Hitchcock et al. 1955–1969, vol. 3). Greene wrote in his description of Viola bakeri that the whole plant was glabrous; he did not mention leaf margins. M. S. Baker (1957) examined the type specimen at UC and apparently was the first to document that its leaf margins are entire. He also noted that under magnification the leaves are ciliate and more or less puberulent. D. M. Fabijan et al. (1987) stated that margins were always entire and ciliate and more or less pubescent throughout. Some collections of V. bakeri from California have leaves with a sharp point or two on the margin, or crenulations on some portion of the basal or cauline blade margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
L. E. McKinney (1992) considered Viola affinis, and much of what botanists had called V. nephrophylla, to be essentially the same taxon. After studying additional specimens, reviewing literature (H. E. Ballard 1994; A. Haines 2001b), and discussions with others (J. Cayouette, H. E. Ballard, A. Haines, pers. comm.), he chose to maintain these as separate taxa. Reports of V. affinis in the Gulf coastal states based on specimens or photographs are usually attributable to V. missouriensis. Viola affinis reportedly hybridizes with V. hirsutula (= V. ×consobrina House), V. cucullata (= V. ×consocia House), V. brittoniana (= V. ×davisii House), V. sororia (= V. ×filicetorum Greene [as species]), V. sagittata var. sagittata (= V. ×hollickii House), and V. nephrophylla (= V. ×subaffinis House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 121. | FNA vol. 6, p. 120. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. bakeri subsp. shastensis, V. nuttallii var. bakeri | V. affinis var. subarctica, V. crenulata, V. sororia subsp. affinis, V. sororia var. affinis, V. subviscosa, V. venustula |
Name authority | Greene: Pittonia 3: 307. (1898) | Leconte: Ann. Lyceum Nat. Hist. New York 2: 138. (1826) |
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