Viola arvensis |
Viola tomentosa |
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European field violet, European field-pansy, field pansy, field violet, pansy, violette des champs, wild pansy |
felt-leaf violet, woolly or felt-leaf or feltleaf violet, woolly violet |
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Habit | Plants annual, caulescent, not stoloniferous, 5–35 cm. | Plants perennial, caulescent, not stoloniferous, 7–10 cm. |
Stems | 1–5, usually erect, sometimes prostrate or decumbent, branched, subglabrous or puberulent, clustered on taproot. |
1–3(–5), prostrate or decumbent to erect, leafy proximally and distally, densely white-tomentose, from usually vertical, subligneous rhizome. |
Leaves | cauline; stipules palmately lobed, middle lobe oblanceolate, obovate, elliptic, or lanceolate, ± equaling leaf blade, proximal lobes dissected, shorter, margins ciliate, apex acute to obtuse; petiole 0.5–2.3 cm, glabrous or puberulent; blade: proximal ovate to ± oblong, distal narrowly or broadly lanceolate, 0.8–3.4 × 0.3–1.9 cm, base attenuate to ± truncate, margins coarsely crenate-serrate, ciliate or eciliate, apex acute to obtuse, surfaces pubescent abaxially, at least on major veins, glabrous adaxially. |
basal and cauline; basal: 1–6; stipules linear to broadly ovate-oblong, margins entire, sometimes with scattered glandular hairs, apex acute to obtuse; petiole 2–6 cm, densely white-tomentose; blade ± oblanceolate to elliptic, 1.5–5 × 1.4–2.1 cm, base attenuate, usually oblique, margins usually entire, rarely crenate distally, ciliate, apex acute to usually obtuse, mucronulate, surfaces densely white-tomentose abaxially, strigose adaxially; cauline similar to basal except: stipules ovate, lanceolate, oblanceolate, or oblong, margins entire or toothed, densely ciliate with white hairs; petiole 1.5–3.5 cm; blade 1.8–4 × 0.6–1.1 cm. |
Peduncles | 2–8 cm, glabrous or pubescent. |
1–4 cm, densely white-tomentose. |
Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 2–4 mm; petals white to pale yellow on both surfaces, upper 2 ± violet, lower 3 with yellow basal area, often violet-veined, lateral 2 bearded, ± equaling or shorter than sepals, lowest with dark yellow area basally, 7–15 mm, spur blue-violet to purple, elongated, 3–5 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins ciliate-tomentose, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 bearded, lowest 6–11 mm, spur yellow, gibbous, 0.5–1.5 mm; style head bearded; cleistogamous flowers absent. |
Capsules | ± spherical, 5–9 mm, glabrous. |
± spherical, 4–5 mm, densely white-tomentose. |
Seeds | brown, 1.5–1.9 mm. |
brown with lighter brown mottling, 2.5–2.8 mm. |
2n | = 34. |
= 12. |
Viola arvensis |
Viola tomentosa |
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Phenology | Flowering May–Jul. | Flowering May–Aug. |
Habitat | Abandoned fields, roadsides, lawns | Dry, gravelly places, open ponderosa, Jeffrey, lodgepole pine forests |
Elevation | 0–3000 m (0–9800 ft) | 1500–2000 m (4900–6600 ft) |
Distribution |
CA; CT; DC; DE; GA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; SPM; Asia (Siberia); Greenland; Europe; Africa [Introduced in North America]
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CA
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Discussion | A. R. Clapham et al. (1987) noted that Viola arvensis is pollinated by insects and is often selfed; T. Marcussen and T. Karlsson (2010) stated that V. arvensis regularly self-pollinates. Roots of V. arvensis have the odor of wintergreen when crushed (A. E. Radford et al. 1968; W. J. Hayden and J. Clough 1990). Viola arvensis is not vegetatively distinguishable from V. tricolor var. tricolor. G. Halliday (pers. comm.) reported that Viola arvensis occurs in southwestern Greenland, where it is introduced, and is ephemeral on St. Pierre and Miquelon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola tomentosa occurs in El Dorado, Nevada, Placer, Plumas, and Sierra counties. M. S. Baker (1949) reported that nearly every leaf axil of V. tomentosa produces a flower bud and that these buds produce chasmogamous flowers late in season instead of cleistogamous flowers, as do other members of the V. nuttallii complex. Viola tomentosa hybridizes with V. purpurea; the hybrids appeared sterile (M. S. Baker 1949). J. Clausen (1964) reported a putative hybrid between V. tomentosa and V. sheltonii from one location in Sierra County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 121. | FNA vol. 6, p. 160. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | Mnemion arvense, V. tricolor var. arvensis | |
Name authority | Murray: Prodr. Stirp. Gott., 73. (1770) | M. S. Baker & J. C. Clausen: Leafl. W. Bot. 5: 142. (1949) |
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