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European field violet, European field-pansy, field pansy, field violet, pansy, violette des champs, wild pansy

alpine marsh violet, marsh violet, northern marsh or marsh or swamp or alpine marsh violet, northern marsh violet, violette des marais

Habit Plants annual, caulescent, not stoloniferous, 5–35 cm. Plants perennial, acaulescent, stoloniferous, 3–21.5 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy.
Stems

1–5, usually erect, sometimes prostrate or decumbent, branched, subglabrous or puberulent, clustered on taproot.

Leaves

cauline;

stipules palmately lobed, middle lobe oblanceolate, obovate, elliptic, or lanceolate, ± equaling leaf blade, proximal lobes dissected, shorter, margins ciliate, apex acute to obtuse;

petiole 0.5–2.3 cm, glabrous or puberulent;

blade: proximal ovate to ± oblong, distal narrowly or broadly lanceolate, 0.8–3.4 × 0.3–1.9 cm, base attenuate to ± truncate, margins coarsely crenate-serrate, ciliate or eciliate, apex acute to obtuse, surfaces pubescent abaxially, at least on major veins, glabrous adaxially.

basal, 2–4, ascending to erect;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1–17 cm, glabrous;

blade unlobed, reniform or ovate to orbiculate, 0.5–6.4 × 0.5–5.5 cm, ± as long as wide, base cordate, margins crenulate, eciliate, apex obtuse to acuminate, surfaces glabrous.

Peduncles

2–8 cm, glabrous or pubescent.

2–20.7 cm, glabrous, bracteoles usually below middle in chasmogamous flowers.

Flowers

sepals lanceolate, margins ciliate or eciliate, auricles 2–4 mm;

petals white to pale yellow on both surfaces, upper 2 ± violet, lower 3 with yellow basal area, often violet-veined, lateral 2 bearded, ± equaling or shorter than sepals, lowest with dark yellow area basally, 7–15 mm, spur blue-violet to purple, elongated, 3–5 mm;

style head bearded; cleistogamous flowers absent.

sepals ovate or lanceolate, margins eciliate, auricles 1–2 mm;

petals lilac, pale blue, or pale violet on both surfaces, sometimes nearly white and tinged with lilac, lowest and sometimes lateral 2 dark violet-veined, lateral 2 bearded or beardless, lowest 8–16 mm, spur usually same color as petals, gibbous, 1–3 mm;

style head beardless; cleistogamous flowers on ascending peduncles.

Capsules

± spherical, 5–9 mm, glabrous.

ellipsoid, 6–10 mm, glabrous.

Seeds

brown, 1.5–1.9 mm.

dark brown, 1–2 mm.

2n

= 34.

= 48.

Viola arvensis

Viola palustris

Phenology Flowering May–Jul. Flowering Apr–Jul.
Habitat Abandoned fields, roadsides, lawns Marshes, swamps, fens, wet forests, stream banks, brushy places, lowlands to subalpine zones
Elevation 0–3000 m (0–9800 ft) 0–1800 m (0–5900 ft)
Distribution
from FNA
CA; CT; DC; DE; GA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; SPM; Asia (Siberia); Greenland; Europe; Africa [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; ME; MT; NH; NV; OR; SD; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

A. R. Clapham et al. (1987) noted that Viola arvensis is pollinated by insects and is often selfed; T. Marcussen and T. Karlsson (2010) stated that V. arvensis regularly self-pollinates. Roots of V. arvensis have the odor of wintergreen when crushed (A. E. Radford et al. 1968; W. J. Hayden and J. Clough 1990). Viola arvensis is not vegetatively distinguishable from V. tricolor var. tricolor.

G. Halliday (pers. comm.) reported that Viola arvensis occurs in southwestern Greenland, where it is introduced, and is ephemeral on St. Pierre and Miquelon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola palustris occurs in small colonies; individual plants are interconnected by stolons. When flowering, usually three or more leaves are present per caudex.

M. Sorsa (1968) noted that in western North America Viola palustris specimens differ from European V. palustris plants and often show variation in character combinations. She stated (pers. comm.) that North American V. palustris is not a uniform entity.

E. Hultén (1968) synonymized Viola palustris with V. epipsila, as did W. J. Cody (2000). G. W. Douglas et al. (1998–2002, vol. 5) recognized both species. We assume that V. palustris does not occur in Alaska; verification is needed. See additional discussion under 18a. V. epipsila var. repens.

M. Sorsa (1968) indicated that Viola palustris var. brevipes has often been confused with V. blanda and that var. brevipes may have arisen from hybridization between a tetraploid and a diploid plant. Listed here in synonymy, var. brevipes may warrant recognition.

Reports of Viola palustris from one location in Coconino County, Arizona, first collected in 1962, likely refer to V. macloskeyi.

Viola palustris hybridizes with V. epipsila var. repens (M. Sorsa 1968; T. Marcussen and T. Karlsson 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 121. FNA vol. 6, p. 142.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms Mnemion arvense, V. tricolor var. arvensis V. palustris subsp. brevipes, V. palustris var. brevipes, V. palustris var. leimonia, V. palustris var. pensylvanica, V. palustris subsp. pubifolia
Name authority Murray: Prodr. Stirp. Gott., 73. (1770) Linnaeus: Sp. Pl. 2: 934. (1753)
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