Viola arvensis
Viola palmata
European field violet, European field-pansy, field pansy, field violet, pansy, violette des champs, wild pansy
early blue violet, three-lobe violet, three-lobed violet, trilobed violet, wood violet
1–5, usually erect, sometimes prostrate or decumbent, branched, subglabrous or puberulent, clustered on taproot.
cauline;
stipules palmately lobed, middle lobe oblanceolate, obovate, elliptic, or lanceolate, ± equaling leaf blade, proximal lobes dissected, shorter, margins ciliate, apex acute to obtuse;
petiole 0.5–2.3 cm, glabrous or puberulent;
blade: proximal ovate to ± oblong, distal narrowly or broadly lanceolate, 0.8–3.4 × 0.3–1.9 cm, base attenuate to ± truncate, margins coarsely crenate-serrate, ciliate or eciliate, apex acute to obtuse, surfaces pubescent abaxially, at least on major veins, glabrous adaxially.
basal, 2–3, ascending to erect;
stipules linear-lanceolate, margins entire, apex acute;
petiole 1–20 cm, glabrous or pubescent;
earliest leaf blades unlobed, mid-season blades 3–9-lobed, lobes sometimes appearing petiolate and sometimes further lobed;
earliest leaf blades reniform to ovate, mid-season blades with middle lobes usually ovate or elliptic to widely obovate, sometimes narrowly elliptic, narrowly ovate, lanceolate, or spatulate to narrowly obovate, lateral lobes elliptic, obdeltate, or spatulate to falcate, smaller lobes similar, 1–14 × 1–10 cm, base truncate to cordate, margins serrate or entire, usually ciliate, apex acute, rounded, blunt, or obtuse, surfaces glabrous or pubescent throughout or along veins.
2–8 cm, glabrous or pubescent.
3–13 cm, glabrous or pubescent.
sepals lanceolate, margins ciliate or eciliate, auricles 2–4 mm;
petals white to pale yellow on both surfaces, upper 2 ± violet, lower 3 with yellow basal area, often violet-veined, lateral 2 bearded, ± equaling or shorter than sepals, lowest with dark yellow area basally, 7–15 mm, spur blue-violet to purple, elongated, 3–5 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm;
petals violet on both surfaces, lower 3 white basally, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, spur sometimes bearded, lowest 15–25 mm, spur white, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
± spherical, 5–9 mm, glabrous.
ellipsoid, 5–15 mm, glabrous.
brown, 1.5–1.9 mm.
beige, mottled to bronze, 1.5–2.5 mm.
= 34.
= 54.
Viola arvensis
Viola palmata
A. R. Clapham et al. (1987) noted that Viola arvensis is pollinated by insects and is often selfed; T. Marcussen and T. Karlsson (2010) stated that V. arvensis regularly self-pollinates. Roots of V. arvensis have the odor of wintergreen when crushed (A. E. Radford et al. 1968; W. J. Hayden and J. Clough 1990). Viola arvensis is not vegetatively distinguishable from V. tricolor var. tricolor.
G. Halliday (pers. comm.) reported that Viola arvensis occurs in southwestern Greenland, where it is introduced, and is ephemeral on St. Pierre and Miquelon.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
N. L. Gil-Ad (1997) presented evidence based on seed coat micromorphology that some plants known as Viola palmata are hybrids between plants with lobed and unlobed leaves and believed the type specimen to be hybrid in origin. He chose not to recognize the name V. palmata, choosing instead to recognize V. triloba. If the type specimen does represent a hybrid, Gil-Ad would be correct in recognizing V. triloba. Because the purity of the type specimen cannot be ascertained, we consider V. palmata the most appropriate name for this taxon. We acknowledge the presence of hybrids between lobed and unlobed plants; such hybrids obscure lines of demarcation. Some herbarium specimens may represent such hybrids; we feel most do not.
L. E. McKinney (1992) described the nomenclatural history of Viola palmata. Homophylly versus heterophylly differentiates V. palmata from V. subsinuata, the homophyllous violet most often called V. palmata. We know that the taxon described by Linnaeus was heterophyllous, with some undivided leaf blades.
Varieties 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Petioles, leaf surfaces, and peduncles usually pubescent, rarely glabrous; apex of middle leaf lobe acute; plants of dry to mesic habitats. | var. palmata |
1. Petioles, leaf surfaces, and peduncles usually glabrous, rarely pubescent; apex of middle leaf lobe rounded, blunt, or obtuse; plants of wet ground. | var. heterophylla |
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