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speedwell, spring speedwell

Cusick's speedwell

Habit Annuals. Perennials.
Stems

erect or ascending, (1–)3–15(–20) cm, at least distally glandular- and eglandular-hairy.

erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy.

Leaves

blade lanceolate to ovate, (4–)6–13 × (2.5–)4–10 mm, 1.3–1.6 times as long as wide, base cuneate, proximal margins coarsely crenate-serrate, distal pinnatifid to subpalmatifid, lobes 3–7, central largest, lateral linear to lanceolate, apex of central lobe obtuse, lateral ± acute, surfaces sparsely glandular-hairy.

blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate.

Racemes

1–3, terminal and axillary, 20–80 mm, (5–)15–40(–60)-flowered, axis eglandular- and glandular-hairy;

bracts proximalmost often 3-fid, others linear-lanceolate, 3–5(–8) mm.

1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy;

bracts lanceolate, 3–7(–11) mm.

Pedicels

erect, (0.4–)1–3 mm, shorter than subtending bract, length 1/3–1/2 times calyx, eglandular- and glandular-hairy.

erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy.

Flowers

calyx lobes (2–)3–5(–7) mm, apex acute, eglandular- and glandular-hairy;

corolla sky to pale blue, 1.5–3 mm diam.;

stamens 0.2–0.6 mm;

style 0.2–0.6 mm, stigma white.

calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy;

corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.;

stamens 4–8 mm;

style (5–)6–9(–10) mm, conspicuously exerted.

Capsules

compressed in cross section, obcordiform, 2.5–3.5 × 3.5–5 mm, apex emarginate, eglandular-hairy.

compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy.

Seeds

8–20(–26), yellowish, ellipsoid, flat, 0.9–1.6 × 0.6–1.3 mm, 0.2 mm thick, smooth to ± rugulose.

28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough.

2n

= 16 (Eurasia).

= 18, 72.

Veronica verna

Veronica cusickii

Phenology Flowering (Mar–)Apr–Jun(–Aug). Flowering Jun–Aug.
Habitat Open pine and oak forests, rocky and sandy steppes, pastures, meadows. Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows.
Elevation 300–2600 m. (1000–8500 ft.) 1500–3200 m. (4900–10500 ft.)
Distribution
from FNA
ID; IN; MA; MI; MN; MT; NY; OR; WA; WI; WY; BC; NB; NS; ON; PE; Eurasia [Introduced in North America]
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[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Specimens of Veronica verna from Alberta have not been verified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 310.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. allenii
Name authority Linnaeus: Sp. Pl. 1: 14. (1753) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878)
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