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speedwell, spring speedwell

bilobed speedwell, sawtooth speedwell, twolobed speedwell

Habit Annuals. Annuals.
Stems

erect or ascending, (1–)3–15(–20) cm, at least distally glandular- and eglandular-hairy.

erect, (7–)10–20(–30) cm, eglandular- and/or glandular-hairy.

Leaves

blade lanceolate to ovate, (4–)6–13 × (2.5–)4–10 mm, 1.3–1.6 times as long as wide, base cuneate, proximal margins coarsely crenate-serrate, distal pinnatifid to subpalmatifid, lobes 3–7, central largest, lateral linear to lanceolate, apex of central lobe obtuse, lateral ± acute, surfaces sparsely glandular-hairy.

blade lanceolate or elliptic, (6–)12–28(–35) × 3–12(–20) mm, base cuneate, margins serrate, teeth 4–7(–12) per side, apex acute, surfaces sparsely glandular-hairy or glabrate.

Racemes

1–3, terminal and axillary, 20–80 mm, (5–)15–40(–60)-flowered, axis eglandular- and glandular-hairy;

bracts proximalmost often 3-fid, others linear-lanceolate, 3–5(–8) mm.

1–8, terminal, usually reduced and replaced by 2 pseudodichotomous racemes with additional axillary racemes, 80–150 mm, (5–)10–25(–35)-flowered, axis eglandular- and/or glandular-hairy;

bracts lanceolate, 9–12 mm, proximals with margins serrate, distals sometimes entire.

Pedicels

erect, (0.4–)1–3 mm, shorter than subtending bract, length 1/3–1/2 times calyx, eglandular- and glandular-hairy.

erect-patent to spreading proximally, deflexed distally, (3–)6–9(–12) mm, shorter than subtending bract, densely glandular- and eglandular-hairy.

Flowers

calyx lobes (2–)3–5(–7) mm, apex acute, eglandular- and glandular-hairy;

corolla sky to pale blue, 1.5–3 mm diam.;

stamens 0.2–0.6 mm;

style 0.2–0.6 mm, stigma white.

calyx lobes (5–)6–8.5(–10) mm, apex acuminate-acute, pairs connate 0.8–1.1 mm, glandular- and eglandular-hairy, rarely sparsely hairy;

corolla mostly intense blue with white center, 4–6 mm diam.;

stamens 2.5 mm;

style (0.7–)1–1.3(–1.7) mm.

Capsules

compressed in cross section, obcordiform, 2.5–3.5 × 3.5–5 mm, apex emarginate, eglandular-hairy.

slightly compressed in cross section, 3.5–5 × (4–)5–8 mm, apex emarginate, sinus angle 30–45(–60)°, densely eglandular- and glandular-hairy, rarely eglandular-hairy.

Seeds

8–20(–26), yellowish, ellipsoid, flat, 0.9–1.6 × 0.6–1.3 mm, 0.2 mm thick, smooth to ± rugulose.

4–10, yellowish brown, oblong to ovoid, cymbiform, 1.5–2.5 × 0.8–1.5 mm, 0.8 mm thick, cristate, dorsally reticulate-verrucate.

2n

= 16 (Eurasia).

= 18, 42 (Asia).

Veronica verna

Veronica argute-serrata

Phenology Flowering (Mar–)Apr–Jun(–Aug). Flowering Apr–Jun.
Habitat Open pine and oak forests, rocky and sandy steppes, pastures, meadows. Bare soils, oak and juniper forests, steppes, alpine meadows, fields, stony slopes, streams, gardens, ruins, ruderal places, calcareous and siliceous soils.
Elevation 300–2600 m. (1000–8500 ft.) (600–)900–3000 m. ((2000–)3000–9800 ft.)
Distribution
from FNA
ID; IN; MA; MI; MN; MT; NY; OR; WA; WI; WY; BC; NB; NS; ON; PE; Eurasia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; IL; KS; MT; NE; NM; NV; NY; OR; UT; WA; WY; BC; Eurasia [Introduced in North America]
[WildflowerSearch map]
Discussion

Specimens of Veronica verna from Alberta have not been verified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 317. FNA vol. 17, p. 321.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. bornmuelleri
Name authority Linnaeus: Sp. Pl. 1: 14. (1753) Regel & Schmalhausen: Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 626. (1878) — (as argute serrata)
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