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undulate speedwell, wavy-leaf water speedwell

trilobed speedwell

Habit Annuals or perennials. Annuals.
Stems

erect or prostrate basally, 10–100 cm, glabrous at least proximally, glandular-hairy distally.

decumbent, (5–)10–30 cm, ± eglandular-hairy.

Leaves

petiole 0–5 mm;

blade elliptic to ovate, sometimes ovate-oblong or linear-lanceolate, rarely lanceolate, 20–50(–100) × 5–20(–25) mm, 2.5–4 times as long as wide, base attenuate, upwards amplexicaul, margins subentire or crenate to serrate or ± undulate, apex acute to obtuse, surfaces glabrous.

blade broadly ovate to suborbiculate, 3(–5)-lobed, central lobe 1/4–1/2 times length of whole leaf, (3–)5–10(–15) × (4–)6–12(–18) mm, ± fleshy, base truncate to subcordate, margins 3(–5)-lobed, apex acute, surfaces sparsely eglandular-hairy.

Racemes

6–25, axillary, 50–220 mm, 10–15 mm diam., 10–100-flowered, axis sparsely glandular-hairy, rarely glabrous;

pedicels 2–6 per cm;

bracts linear to lanceolate, 3–6 mm, apex acute.

1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis sparsely eglandular-hairy;

bracts broadly ovate to suborbiculate, 3(–5)-lobed, (3–)5–10(–15) mm.

Pedicels

patent, 3–5 mm, equal to subtending bract, sparsely glandular-hairy.

patent, deflexed in fruit, (2–)4–8(–10) mm, shorter than subtending bract, length 1–2 times calyx, glabrous abaxially, hairy adaxially.

Flowers

calyx lobes 1.5–2(–3) mm, apex acute, sparsely glandular-hairy, rarely glabrous;

corolla pale blue or pale lilac, rarely white, 2.5–5 mm diam.;

stamens 2 mm;

style 0.9–2 mm.

calyx lobes 4–5(–6) mm, apex acute, puberulent on abaxial surface, rarely glabrous, densely ciliate with 35–60 per side;

corolla deep blue with white center, 3–5 mm diam.;

stamens dark blue, length unknown;

style 0.5–0.9(–1.1) mm.

Capsules

slightly compressed in cross section, obcordiform to globular, 2–3 × 2–3 mm, apex rounded or ± emarginate, sparsely glandular-hairy.

± round in cross section, ovoid, 2.5–3(–3.5) × 4–5 mm, apex ± emarginate, glabrous.

Seeds

30–40, ochre, broadly ellipsoid, ± flat, convex on both sides, 0.5–0.6 × 0.4–0.5 mm, thickness and texture unknown.

(1–)4, dull yellowish brown, ellipsoid, often elongate-cuboidal, deeply urn-shaped (elaiosome inside), 2.2–2.8 × 1.5–2.2 mm, 1–1.4 mm thick, cristate to strongly rugose at margins.

2n

= 54 (Asia).

= 18 (Eurasia).

Veronica undulata

Veronica triloba

Phenology Flowering (Mar–)Apr–Sep. Flowering (Feb–)Mar–May(–Jun).
Habitat Disturbed, wet places, ditches, or swamps. Disturbed sites.
Elevation 0–200 m. (0–700 ft.) 100–200 m. (300–700 ft.)
Distribution
from FNA
AL; OR; WA; Asia [Introduced in North America]
[BONAP county map]
from FNA
CA; OH; Europe (Balkans); Asia (Turkey) [Introduced in North America]
Discussion

In the flora area, Veronica undulata was introduced near ports (Mobile, Alabama, and Portland, Oregon) before 1900 via ship ballast from trade with Asia, did not spread much, and may not have persisted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Veronica triloba is known in the flora area from three recent collections in California and Ohio; it is possibly more widespread than reported here. It is closely related to V. hederifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 314. FNA vol. 17, p. 316.
Parent taxa Plantaginaceae > Veronica Plantaginaceae > Veronica
Sibling taxa
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triloba, V. triphyllos, V. verna, V. wormskjoldii
V. agrestis, V. alpina, V. americana, V. anagallis-aquatica, V. argute-serrata, V. arvensis, V. beccabunga, V. biloba, V. catenata, V. chamaedrys, V. copelandii, V. cusickii, V. cymbalaria, V. dillenii, V. filiformis, V. fruticans, V. grandiflora, V. hederifolia, V. longifolia, V. officinalis, V. peregrina, V. persica, V. polita, V. scutellata, V. serpyllifolia, V. spicata, V. stelleri, V. sublobata, V. teucrium, V. triphyllos, V. undulata, V. verna, V. wormskjoldii
Synonyms V. hederifolia var. triloba
Name authority Wallich: in W. Roxburgh, Fl. Ind. 1: 147. (1820) (Opiz) Opiz: Naturalientausch 11: 467. (1826)
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