Veronica triloba |
Veronica cusickii |
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trilobed speedwell |
Cusick's speedwell |
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Habit | Annuals. | Perennials. |
Stems | decumbent, (5–)10–30 cm, ± eglandular-hairy. |
erect, unbranched or sparsely branched at base, (5–)10–15(–30) cm, finely glandular-hairy. |
Leaves | blade broadly ovate to suborbiculate, 3(–5)-lobed, central lobe 1/4–1/2 times length of whole leaf, (3–)5–10(–15) × (4–)6–12(–18) mm, ± fleshy, base truncate to subcordate, margins 3(–5)-lobed, apex acute, surfaces sparsely eglandular-hairy. |
blade elliptic-ovate, (5–)10–25 × 5–10 mm, base cuneate, margins entire or subentire, apex short-acuminate, surfaces glabrous or glabrate. |
Racemes | 1–10, terminal, (50–)100–300 mm, 5–20-flowered, axis sparsely eglandular-hairy; bracts broadly ovate to suborbiculate, 3(–5)-lobed, (3–)5–10(–15) mm. |
1, terminal, (15–)40–80(–130) mm, (4–)8–30(–40)-flowered, axis finely glandular-hairy; bracts lanceolate, 3–7(–11) mm. |
Pedicels | patent, deflexed in fruit, (2–)4–8(–10) mm, shorter than subtending bract, length 1–2 times calyx, glabrous abaxially, hairy adaxially. |
erect, 3–9 mm, equal to or ± longer than subtending bract, finely glandular-hairy. |
Flowers | calyx lobes 4–5(–6) mm, apex acute, puberulent on abaxial surface, rarely glabrous, densely ciliate with 35–60 per side; corolla deep blue with white center, 3–5 mm diam.; stamens dark blue, length unknown; style 0.5–0.9(–1.1) mm. |
calyx lobes 2–3 mm, apex acute to obtuse, finely glandular-hairy; corolla deep blue-violet, rarely white, rotate, (8–)10–13 mm diam.; stamens 4–8 mm; style (5–)6–9(–10) mm, conspicuously exerted. |
Capsules | ± round in cross section, ovoid, 2.5–3(–3.5) × 4–5 mm, apex ± emarginate, glabrous. |
compressed in cross section, oblong-obovoid, 4–6 × 3.5–5.5 mm, ca. as long as wide, apex emarginate, finely glandular-hairy. |
Seeds | (1–)4, dull yellowish brown, ellipsoid, often elongate-cuboidal, deeply urn-shaped (elaiosome inside), 2.2–2.8 × 1.5–2.2 mm, 1–1.4 mm thick, cristate to strongly rugose at margins. |
28–32, straw colored or dark brown, ovoid, flat, 0.6 × 0.4 mm, 0.1 mm thick, rough. |
2n | = 18 (Eurasia). |
= 18, 72. |
Veronica triloba |
Veronica cusickii |
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Phenology | Flowering (Feb–)Mar–May(–Jun). | Flowering Jun–Aug. |
Habitat | Disturbed sites. | Gravelly soils, openings in coniferous forests and in subalpine and alpine meadows. |
Elevation | 100–200 m. (300–700 ft.) | 1500–3200 m. (4900–10500 ft.) |
Distribution |
CA; OH; Europe (Balkans); Asia (Turkey) [Introduced in North America] |
CA; ID; MT; OR; WA; BC
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Discussion | Veronica triloba is known in the flora area from three recent collections in California and Ohio; it is possibly more widespread than reported here. It is closely related to V. hederifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Veronica cusickii is sister to V. copelandii and closely related to V. wormskjoldii subsp. nutans (D. C. Albach et al. 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 316. | FNA vol. 17, p. 310. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | V. hederifolia var. triloba | V. allenii |
Name authority | (Opiz) Opiz: Naturalientausch 11: 467. (1826) | A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 288. (1878) |
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